ML tree based on the 28S + COI dataset (1,075 bp in length). Ultrafast bootstrap (UFB), SH-aLRT and posterior probability (pp) values are given beside the nodes. The values were omitted when UFB < 95, SH-aLRT < 80 and pp < 0.90. Tips are labelled with specimen ID. Tokyo, Tokyo Metropolis; Yakushima, Yakushima Island; Amami, Amami-Oshima Island; Okinawa, Okinawa-Hontou Island; Irabu, Irabu–jima Island; N. Taiwan, Northern Taiwan; C. Taiwan, Central Taiwan; E. Taiwan, Eastern Taiwan; S. Taiwan, Southern Taiwan.