Tandonia rustica. A–C sequenced specimen NMBE 568973, Rovio, Sovaglia A preserved animal B situs of genitalia C distal genitalia showing anatomical details D penis papillae of sequenced specimen NMBE 568978, Carona E–G sequenced specimen NMBE 564708, Goldswil E situs of genitalia F distorted epiphallus G details of lumina of vagina, atrium and penis, arrows pointing at the “twin papillae” (tp) H–J spermathophore of specimen NMBE 571414, Orselina H genital pore with piece of spermatophore I, J largest part of the three pieces J detail of surface covering spines. Abbreviations: a = atrium; bc = bursa copulatrix; ep = epiphallus; fod = free oviduct; p = penis; ped = pedunculus; pp = penial papilla; spo = spermoviduct; tp = “twin papillae”; v = vagina; vd = vas deferens; vg = vaginal gland.