The distribution of Nebria (Sadonebria) spp. in the Japanese Archipelago; N. chinensis Bates, which is distantly related to the remaining species, and samples that are not identified by genital morphology are not included [modified from Sasakawa and Toki (2011)]. 1 N. sadona sadona Bates 2 N. saeviens Bates 3 N. asahina Sasakawa 4 N. nasuensis Sasakawa 5 N. sadona leechi Bates 6 N. quinquelobata sp. n. 7 N. chichibuensis Sasakawa 8 N. yatsugatakensis sp. n. 9 N. kiso Sasakawa 10 N. mikawa Sasakawa 11 N. sadona ohdaiensis Nakane 12 N. tenuicaulis Sasakawa & Kubota 13 N. jakuchisana Sasakawa 14 N. shikokuensis Sasakawa; 15: N. trifida Sasakawa.