Strong differentiation between amphibian communities on two adjacent mountains in the Upper Rio Pastaza watershed of Ecuador, with descriptions of two new species of terrestrial frogs

Sample points in the Naturetrek-Vizcaya Reserve (black triangles) and the Machay Reserve (black circles). The white triangle is the type locality of Pristimantis maryanneae sp. nov. and the white circle is the type locality of Pristimantis burtoniorum sp. nov. The grey shaded areas are the Reserves of the EcoMinga Foundation within the Llanganates-Sangay Corridor and the green shaded areas represent Llanganates National Park (north) and Sangay National Park (south).