Bringing order to a complex system: phenotypic and genotypic evidence contribute to the taxonomy of Tityus (Scorpiones, Buthidae) and support the description of a new species

Tityus C. L. Koch, 1836, telotarsi IV, showing ventral macrosetae A, C, E, G, I, K white light B, D, F, H, J, L UV light A, B Tityus (Tityus) brazilae Lourenço & Eickstedt, 1984 (type II) (MZSP 75619) C, D Tityus (Archaeotityus) clathratus C. L. Koch, 1844 (type I) (MZSP 31468) E, F Tityus (Atreus) forcipula (Gervais, 1843) (type II) (MZSP) G, H Tityus (Atreus) obscurus Gervais, 1843 (type I) (MNRJ 07610) I, J Tityus (Tityus) serrulatus Lutz & Mello, 1922 (type II) (MZSP 28205) K, L Tityus (Tityus) spelaeus sp. nov. (MZSP 74633) (type II). Observations = telotarsi I–IV ventral setae distribution: Type I = tuft of irregularly distributed setae. Type II = two discrete ventrosubmedian rows of setae. Scale bars: 500 μm.