Two new species of the genus Camptoscaphiella from Yunnan, China (Araneae, Oonopidae)

Abstract Two new species of the genus Camptoscaphiella Caporiacco, 1934 are described from Yunnan, China, i.e., C. changxu Tong & Li, sp. nov. (♂♀) and C. linyejiei Tong & Li, sp. nov. (♂♀). Camptoscaphiella changxu Tong & Li, sp. nov. is characterized by the long, strongly-curved setae between male paturons, which is unknown in other oonopids and in any other spiders. Camptoscaphiella linyejiei Tong & Li, sp. nov. is the third blind species of this genus in the world. Morphological descriptions and photographic illustrations of the two new species are given.


Introduction
Camptoscaphiella Caporiacco, 1934, is a small genus of oonopid spiders that currently contains 18 species (Li 2020;WSC 2021). It is distributed in montane tropical and subtropical regions in Asia, mostly within the Himalayan Plateau (Baehr and Ubick 2010). This genus can be recognized by the remarkable morphology of the male palp, which has an extremely large, club-shaped palpal patella and a bulb that is not fused with the cymbium, and the first two pairs of legs which have extremely long spines with the tibiae bearing 4 pairs of spines and the metatarsi bearing 2 pairs of spines (Baehr and Harvey 2013).
Camptoscaphiella is still poorly studied. Currently three species of this genus are recorded in Yunnan, China, i.e., C. paquini Ubick, 2010, C. sinensis Deeleman-Reinhold, 1995and C. tuberans Tong & Li, 2007, and only one species, C. schwendingeri Baehr, 2010is recorded in Thailand (Deeleman-Reinhold 1995Tong and Li 2007;Baehr and Ubick 2010). There is no species recorded in the adjacent areas of south of Yunnan, i.e., Laos, Myanmar and Vietnam. In this paper two new Camptoscaphiella species, C. changxu Tong & Li, sp. nov. and C. linyejiei Tong & Li, sp. nov. collected from Yunnan, are described and illustrated.

Materials and methods
The specimens were examined using a Leica M205C stereomicroscope. Details were studied under an Olympus BX51 compound microscope. Photos were made with a Canon EOS 550D zoom digital camera (18 megapixels) mounted on an Olympus BX51 compound microscope. Vulvae were cleared in lactic acid. For scanning electron microscopy (SEM), specimens were air-dried, sputter-coated using IXRF SYSTEMS, and imaged with a Hitachi TM3030 SEM. All measurements were taken using an Olympus BX51 compound microscope and are in millimeters. The type material is deposited in Shenyang Normal University (SYNU) in Shenyang, China.

Family Oonopidae Simon, 1890
Genus Camptoscaphiella Caporiacco, 1934 Type species. Camptoscaphiella fulva Caporiacco, 1934, by monotypy. Diagnosis. Males of this genus can easily be separated from all other oonopid genera by the heart-shaped sternum with conical projection on the anterolateral corners, spination of the first two legs (tibia I and II with four pairs of long spines, and metatarsus I and II with two pairs of long spines); the extremely large, club-shaped palpal patella; and a cymbium that is not fused with the bulb. Females of this genus are similar to those of Ischnothyreus Simon, 1893, but can be separated by lacking the distinct, darkly sclerotized, strongly winding duct and uniquely shaped atrium (revised from Baehr and Harvey 2013).
Comments. The male chelicerae are unmodified in species considered to belong to the genus (Baehr and Ubick 2010). The long, strongly-curved setae between male cheliceral paturon of this species are unique in this genus, even in the entire order.
Distribution. Known only from the type locality. Diagnosis. This new species is similar to C. sinensis, but can be distinguished by the flat carapace (Fig. 4F), the presence of dorsal and ventral abdominal scuta (Fig. 4A, C, E), the unmodified cymbium (Fig. 5E), and the short processes of tip of the bulb (Fig. 5G, H). Camptoscaphiella sinensis has the highest point of carapace at posterior 2/3, the abdomen lacking scuta, the tip of the cymbium with a pair of enlarged tubular setae, and the tip of bulb with elongated processes (Deeleman-Reinhold 1995: figs 1-5).
Etymology. The specific name is named after Mr Yejie Lin, the collector of the type specimens; noun (name) in genitive case.
Distribution. Known only from the type locality.