A new genus and two new species of oonopid spiders from Tibet, China (Araneae, Oonopidae)

Abstract A new genus, Paramolotra Tong & Li, gen. nov., including two new species, Paramolotra pome Tong & Li, sp. nov. (♂♀) and Paramolotra metok Tong & Li, sp. nov. (♂♀), is described from Tibet, China. Morphological descriptions and photographic illustrations of the two new species are given.

The oonopid spiders of Tibet have been poorly studied. Hitherto, only two species, Gamasomorpha linzhiensis Hu, 2001 and Ischnothyreus linzhiensis Hu, 2001, have been recorded from Tibet (Hu 2001). In this paper, a new genus with two new species are proposed from material collected from Tibet are described and illustrated.

Materials and methods
The specimens were examined using a Leica M205C stereomicroscope. Details were studied under an Olympus BX51 compound microscope. Photos were made with a Canon EOS 550D zoom digital camera (18 megapixels) mounted on an Olympus BX51 compound microscope. Vulvae were cleared in lactic acid. For scanning electron microscopy (SEM), specimens were air-dried, sputter coated using IXRF SYSTEMS, and imaged with a Hitachi TM3030 SEM. All measurements were taken using an Olympus BX51 compound microscope and are in millimeters. The type material is deposited in Shenyang Normal University (SYNU) in Shenyang, China (curator: Yanfeng Tong).
Etymology. The generic name refers to the similarities of this genus with Promolotra Tong & Li and is feminine in gender.
Diagnosis. Paramolotra gen. nov. resembles Promolotra Tong & Li, 2020 in having the heavily sclerotized dorsal and ventral abdominal scuta, the long spines on legs I and II, the cone-shaped protuberance on anterior face of male chelicerae, and the completely fused bulb and cymbium, but can be distinguished by the embolar region which have several protrusions that distinctly extend beyond the tip of the cymbiobulbus (Figs 2, 5, 7A-D), the indented labium (Fig. 8A, G) of both sexes, and the stick-shaped anterior arm of T-shaped sclerite (tsc) of endogyne (Fig. 9B, D). The embolar region of Promolotra consists of brush-like structures and 3 broad lobes, which barely extends beyond the tip of the cymbiobulbus (Tong and Li 2020: figs 1H-J, 2, 5H-J, 6), the labium deeply incised (Tong and Li 2020: fig. 3E, 7E), the endogyne consists of canopy-shaped anterior arm of T-shaped sclerite (tsc) and horseshoe-shaped sclerite (Tong and Li 2020: fig. 4G).
Female (paratype, SYNU-438). As in male except as noted. Body: habitus as in Fig. 3A Etymology. The specific name is a noun in apposition taken from the type locality. Distribution. Known only from the type locality (Fig. 10).          Diagnosis. This new species is similar to Paramolotra pome sp. nov., but can be distinguished by the long, curved hook like anterior protrusion (ap) of embolar region (Fig. 5), and the very short anterior arm of T-shaped sclerite of endogyne (Fig. 9D). Paramolotra pome sp. nov. males have the narrow anterior protrusion (ap) of embolar region (Fig. 2), and females have very long anterior arm of T-shaped sclerite of endogyne (Fig. 9B).
Etymology. The specific name is a noun in apposition taken from the type locality. Distribution. Known only from the type locality (Fig. 10).

Discussion
The homologies in the genitalia of males and females of Paramolotra gen. nov. are unclear. Paramolotra gen. nov. are very similar to Promolotra in the fused palpal bulb and cymbium and the somatic characters. The embolar region of Promolotra has 3 leaf-like, wrinkled texture, and nearly translucent lobes (Tong and Li 2020: figs 1H-J, 5H-J), which is different from the sclerotized protrusions of the embolar region ( Fig. 7A-D) of Paramolotra gen. nov. The endogyne of Promolotra has a horseshoe-shaped sclerite (Tong and Li 2020: fig. 4G), which is lacking in the new genus. However, the T-shaped sclerite and the tube-like structure of endogyne (Tong and Li 2020: figs 4G; 9B, D) are quite homogeneous. As for the other Asian oonopid genera, i.e., Kachinia Tong &Li, 2018, andVientianea Tong &Li, 2013, also have heavily sclerotized abdominal scuta and leg spines. Paramolotra gen. nov. are quite different from both genera. The genus Kachinia differs from Paramolotra gen. nov. by the heavily sclerotized and darkened palps of males (Tong et al. 2018: figs 1I-K, 4I-K), and the tube-like posterior receptacle of endogyne (Tong et al. 2018: figs 3I, J, 6I, J). The genus Vientianea differs from Paramolotra gen. nov. by the enlarged male palpal patella (Tong and Li 2013: figs 24, 25), and the medially stick-shaped sclerite and the strongly curved circular sclerite of endogyne (Tong and Li 2013: figs 23, 33). So, Paramolotra gen. nov. and Promolotra maybe represent a different genus group in Asia.