Boxplot of the standardized catch of epigaeic beetles at seven locations in aspen-dominated mixedwood forests in north-central Alberta, 1992–93. Locations are distributed across four Natural Subregions (SR) and two Natural Regions (NR) A Carabidae B Staphylinidae. Locations with the same letter situated above the bar are not significantly different at α = 0.05 (post hoc Tukey’s tests). For locations where there was significant interannual variation, the results are indicated above the bar for that location.