A review of the spider-attacking Polysphinctadizardi species-group (Hymenoptera, Ichneumonidae, Pimplinae), with descriptions of seven new species from South America

Abstract The Neotropical Polysphinctadizardi species-group is revised. We describe seven new species from South America: P.bonitasp. nov., P.cosnipatasp. nov., P.incasp. nov., P.macroepomiasp. nov., P.organensissp. nov., P.pichinchasp. nov., and P.teresasp. nov. In addition, we provide a diagnosis and an identification key to all species of the group.


Introduction
UFMG Universidade Federal de Minas Gerais, Belo Horizonte, Minas Gerais, Brazil; ZMUT Biodiversity Unit, Zoological Museum of the University of Turku, Turku, Finland.
The morphological terminology follows Broad et al. (2018) and style of the descriptions follow those of . However, we add two new characters to the descriptions: the shape of the tarsal claws and the shape of the pronotal shelf. We also add the following proportions to the descriptions: (a) margin of gena/length of eye; (b) length of the epomia/length of the proximal mandibular width; and (c) length/ posterior width of tergite II.
The measures and proportions between the structures are given as the value of the holotypes or paratypes [in brackets], followed by the minimum and maximum number of variations. The [brackets] were also used to add, supplement or correct information on the specimen labels.
Specimens were examined using OLYMPUS SZ61 and SZX10 (at ZMUT) and the ZEISS Stemi 2000 (at INPA) stereomicroscopes. Measurements were obtained using millimetric oculars attached to the stereomicroscope, calibrated with a precision ruler. Digital images were taken using a CANON DS126461 digital camera attached to an OLYMPUS SZX16 stereomicroscope and combined by using the software Zerene Stacker (v. 1.04 Build T201706041920) (at ZMUT) and a LEICA DMC4500 digital camera attached to a LEICA M205A stereomicroscope and combined by using the software Helicon Focus v. 5.3 Pro. (at INPA).
The distributional maps were created using SimpleMappr online software (Shorthouse 2010).

Taxonomy
The Polysphincta dizardi species-group Diagnosis. The P. dizardi species-group can be distinguished from all other speciesgroups of the genus by the combination of two characters: (1) pronotum with a strong shelf-like projection mediodorsally and (2) submetapleural carina absent.
Remarks. According to  and our new discoveries, this species group is known to occur only in the Neotropical region.
Key to the species of the P. dizardi species-group [Obs. Only the males of P. shabui Gauld, P. sinearanea Pádua, and P. organensis sp. nov Metasoma orange, with posterior margins of tergites II-IV narrowly black, posterior half of tergite V black, and tergites VI+ black (  shelf-like projection, in dorsal view, with the apex bilobed, and in lateral view, with anterolateral part posteriorly rounded and weakly decurved; mesoscutum more or less robust, in dorsal view, smooth and polished, with notauli weakly impressed anteriorly; scutellum convex, not laterally carinate; mesopleuron highly polished, virtually impunctate; epicnemial carina reaching almost to the level of the lower corner of pronotum; epicnemium with a vestigial vertical carina near lower corner of pronotum; metapleuron convex, smooth and polished, with sparse, fine bristles evenly spaced, without a discernible submetapleural carina. Propodeum mediodorsally smooth and polished, with longitudinal carinae present only posteriorly and with scattered fine bristles. Fore wing length about [8.0] mm; 1cu-a interstitial relative to M&RS; base of 1m-cu&M separated from CU by about length of 2cu-a; hind wing with distal abscissa of CU present and complete, well pigmented; first abscissa of RS subequal to rs-m. Tarsal claw with proximal lobe quadrangular, with claw apex slightly overtaking the distal margin of lobe. Metasoma.  Color. Head black except posterior 0.8 of clypeus yellowish; antennae brownish with scape and pedicel ventrally yellowish; mouthparts whitish, except apex of mandible blackish. Mesosoma orange. Metasoma with tergite I orange with posterior margin black, tergites II-IV brownish with anterior and anterolateral margins whitish and posterior margin black, and tergites V+ brownish. Fore and mid leg orange, hind leg brownish, except coxa orange and a medium inner and outer region whitish. Wings are hyaline, slightly infuscate, pterostigma brown. Ovipositor brown, with posterior and anterior part whitish. Male Distribution. Brazil (Fig. 13). Biological note. Host unknown. Etymology. The specific name (in apposition) refers to the type locality of this species, RPPN Serra Bonita, Bahia state, Brazil, and also to the beauty of this new species.
Remarks. Polysphincta bonita sp. nov. closely resembles P. organensis sp. nov. mainly by the coloration, with mesosoma entirely orange and metasoma brownish with tergites II-IV or II-VI whitish in anterior and anterolateral margins. It clearly differs from P. organensis sp. nov. by having ovipositor 1.0 times as long as hind tibia and fore and mid leg orange, hind leg brownish, except coxa orange and a medium inner and outer region whitish (ovipositor >1.2 times as long as hind tibia and fore leg orange, mid leg orange with coxa, trochanter and trochantellus whitish and tarsus brownish, hind leg whitish with coxa inner region, trochanter proximally, trochantellus distal, femur proximally and distally, tibia proximally and distally, first tarsal segment distally, and remaining tarsal segments entirely blackish brown in P. organensis sp. nov.).  dorsal view, bilobed, subquadrangular, and in lateral view, slender, with anterolateral corners weakly decurved; mesoscutum more or less robust, in dorsal view, smooth and polished, with notauli weakly impressed anteriorly; scutellum convex, not laterally carinate; mesopleuron highly polished, virtually impunctate; epicnemial carina reaching almost to level of lower corner of pronotum; epicnemium with vestigial vertical carina near lower corner of pronotum; metapleuron convex, smooth and polished, with sparse, fine bristles evenly spaced, without discernible submetapleural carina.

Polysphincta cosnipata
Propodeum mediodorsally smooth and polished, with longitudinal carinae present only posteriorly and with scattered fine bristles. Fore wing length about [7.0] mm; 1cua more or less interstitial relative to M&RS; base of 1m-cu&M separated from CU by about length of 2cu-a; hind wing with distal abscissa of CU present and complete but weakly pigmented; first abscissa of RS subequal to rs-m. Tarsal claw with proximal lobe quadrangular, with claw apex slightly overtaking distal margin of lobe. Metasoma. Tergite I about [1.5] times as long as posteriorly broad, dorsally with lateromedian longitudinal carinae only discernible at extreme anterior part; sternite I with weak swelling near hind rim, and with weak median longitudinal ridge anteriorly; tergite II about [1.5] times as long as posteriorly broad, highly polished, at most with only fine setiferous punctures laterally; tergite III about [1.3] times as long as posteriorly broad, highly polished, at most with only fine setiferous punctures; subgenital plate subquadrate. Ovipositor slightly slender, [1.2] times as long as hind tibia, posteriorly evenly tapered to sharp point.
Color. Head black except 0.8 of clypeus yellowish; antennae brownish with scape and pedicel ventrally whitish; mouthparts whitish, except apex of mandible brownish. Mesosoma orange with metapleuron and propodeum brown. Metasoma brownish with anterior and anterolateral margins of tergites III-V whitish. Fore leg orange, mid leg orange with coxa, trochanter and trochantellus whitish and tarsus distally brownish, hind leg whitish with coxa inner region, trochanter proximal, trochantellus distally, femur proximally and distally, tibia proximally and distally, first tarsal segment distally, and remaining tarsal segments entirely blackish brown. Wings hyaline, pterostigma brown. Ovipositor brown, with posterior and anterior parts whitish.
Remarks. Polysphincta cosnipata sp. nov. closely resembles P. dizardi Gauld, 1991 and P. macroepomia sp. nov. mainly by coloration, with mesosoma orange and propodeum blackish or brownish. However, it differs from P. dizardi by having pronotum orange and metasomal tergites II-VI with anterior and anterolateral margins whitish (anterior part of pronotum brownish and metasomal tergites entirely darkish brown in P. dizardi), and from P. macroepomia sp. nov. by having epomia absent (present in P. macroepomia sp. nov.).  Comments. Additional characters to the original description (♀) are as follows: body about [7.7-8.3] 7.5-8.5; head in dorsal view with margin of the gena convex behind the eyes, and its margin [0.5-0.55] 0.45-0.55 times length of eye; shelf-like projection, in dorsal view, more or less developed anterolaterally, apex very weakly bilobed, and in lateral view, with anterolateral part of apex rounded and very weakly decurved; mesoscutum robust, in dorsal view; tarsal claw with proximal lobe quadrangular, with claw apex slightly overtaking distal margin of lobe.

Polysphincta dizardi Gauld, 1991
Distribution. Costa Rica (Fig. 11). shelf-like projection, in dorsal view, more or less bilobed, subquadrangular, and in lateral view, slender with anterolateral corners weakly decurved; mesoscutum more or less slender, in dorsal view, smooth and polished, with notauli weakly impressed anteriorly; scutellum convex, not laterally carinate; mesopleuron highly polished, virtually impunctate; epicnemial carina reaching almost level of lower corner of pronotum; epicnemium with vestigial vertical carina near lower corner of pronotum; metapleuron convex, smooth and polished, with few sparse, fine bristles evenly spaced, without discernible submetapleural carina. Propodeum mediodorsally smooth and polished, with longitudinal carinae present only posteriorly and laterally with scattered fine bristles. Fore wing length [10.0] 10.0-11.0 mm; 1cu-a interstitial relative to M&RS; base of 1m-cu&M separated from CU by about length of 2cu-a; hind wing with distal abscissa of CU present and complete but weakly pigmented; first abscissa of RS subequal to rs-m. Tarsal claw with proximal lobe quadrangular, with claw apex slightly overtaking the distal margin of lobe. Color. Head black except 0.8 distal of clypeus yellowish; antennae brown; mouthparts pale, except apex of mandible brownish. Mesosoma orange, except posterior carinae of propodeum darkish brown. Metasoma orange, with posterior margins of tergites II-V narrowly black, tergites VI+ black. Fore leg orange, mid leg orange with tarsus brownish, hind leg darkish brown, with median region of tibia pale. Wings yellowish hyaline with apex weakly blackish, pterostigma yellow. Ovipositor darkish brown, with posterior and anterior parts pale.
Variation. Some specimens present hind leg entirely darkish brown; metasoma orange with posterior margins of tergites II-V narrowly black only laterally and tergite VI orange with posterior margin black.
Etymology. This species is named in honour of the Andean Inca empire. Remarks. Polysphincta inca sp. nov. closely resembles A. sinearanea Pádua, 2018 and P. pichincha sp. nov. mainly by color pattern, body orange with last metasomal tergites black. It differs from P. sinearanea by having epomia absent (present in P. sinearanea), and from P. pichincha sp. nov. by having malar space > 0.6 times as long as proximal mandibular width (malar space 0.4 times as long as proximal mandibular width in P. pichincha sp. nov.). (1) epomia present, 1.5 times length of proximal mandibular width (Fig. 5B); (2) malar space 0.6 times as long as proximal mandibular width (Fig. 5B); (3) fore wing vein 1cu-a interstitial relative to M&RS (Fig. 5A); (4) mesosoma orange, except metapleuron and propodeum darkish brown (Fig. 5A); (5) fore wing hyaline (Fig. 5A); (6) hind leg whitish with spot in proximal region of coxa, base of trochanter, longitudinal spot in 1] times as broad as high, weakly convex centrally, polished, with fine sparse setiferous punctures; head in dorsal view with margin of gena very weakly convex behind the eyes, and its margin about [0.6] times length of eye; ocelli moderately large, the lateral ones separated from compound eyes by [1.0] times their own maximum diameter. Mesosoma. Pronotum with epomia distinct, about [1.5] times length of proximal mandibular width; shelf-like projection, in dorsal view, more or less straight, broader than long, and in lateral view, slender and slightly decurved in apex; mesoscutum robust, in dorsal view, smooth and polished, with notauli weakly impressed anteriorly; scutellum convex, not laterally carinate; mesopleuron highly polished, virtually impunctate; epicnemial carina reaching almost the level of lower corner of pronotum; epicnemium with vestigial vertical carina near lower corner of pronotum; metapleuron weakly convex, smooth and polished, with few sparse fine bristles, without discernible submetapleural carina. Propodeum mediodorsally smooth and polished, with longitudinal carinae present only posteriorly and with scattered fine bristles. Fore wing length [6.0] mm; 1cu-a interstitial relative to M&RS; base of 1m-cu&M separated from CU by about length of 2cu-a; hind wing with distal abscissa of CU present and complete but weakly pigmented; first abscissa of RS subequal to rs-m. Tarsal claw with proximal lobe quadrangular, with claw apex slightly overtaking distal margin of lobe. Metasoma. Tergite I about [1.4] times as long as posteriorly broad, dorsally with lateromedian longitudinal carinae only discernible at extreme anterior part; sternite I with weak swelling near hind rim, and with weak median longitudinal ridge anteriorly; tergite II [1.3] times as long as posteriorly broad, highly polished, at most with only fine setiferous punctures laterally; tergite III about [1.2] times as long as posteriorly broad, highly polished, at most with only fine setiferous punctures; subgenital plate subquadrate. Ovipositor slightly slender, [1.7] times as long as hind tibia, posteriorly evenly tapered to sharp point.

Polysphincta macroepomia
Color. Head darkish brown except clypeus brownish; antennae brown; mouthparts white, except apex of mandible black. Mesosoma orange, except metapleuron and propodeum darkish brown. Metasoma entirely darkish brown, with posterior margins of tergites II-V narrowly black. Legs whitish, fore leg with femur, tibia and tarsus weakly yellowish; mid leg with femur and tibia and tarsus weakly yellowish, except final distal of tarsus brownish; hind leg with spot in proximal region of coxa, base of trochanter, longitudinal spot in subdistal region of inner and outer margin of femur, distal part of tibia and final distal of tarsus brownish. Wings hyaline, pterostigma brown. Ovipositor brown, with posterior portion whitish.
Male. Unknown. Distribution. Peru (Fig. 13). Biological notes. Host unknown. Etymology. The specific name refers to the long epomia, main characteristic of this species.
Description. Female. Body [8.5] 7.0-8.5 mm. Head. Clypeus weakly convex, posterior margin thin and straight centrally; malar space [0.4] times as long as proximal mandibular width; lower face about [0.9] 0.9-1.0 times as broad as high, weakly convex centrally, polished, with fine sparse setiferous punctures; head in dorsal view with margin of gena weakly convex behind the eyes, and its margin about [0.5] 0.45-0.5 times length of eye; ocelli moderately large, lateral ones separated from compound eyes by about [0.9] 0.9-1.0 times their own maximum diameter. Mesosoma. Pronotum without epomia; shelf-like projection, in dorsal view, more or less bilobed, subquadrangular, and in lateral view, slender and with anterolateral corners weakly decurved; mesoscutum robust, in dorsal view, smooth and polished, with notauli weakly impressed anteriorly; scutellum convex, not laterally carinate; mesopleuron highly polished, virtually impunctate; epicnemial carina reaching almost level of lower corner of pronotum; epicnemium with vestigial vertical carina near lower corner of pronotum; metapleuron convex, smooth and polished, with few sparse, fine bristles evenly spaced, without discernible submetapleural carina. Propodeum mediodorsally smooth and polished, with longitudinal carinae present only posteriorly and with scattered fine bristles. Fore wing length about [7.0] 5.0-7.0 mm; 1cu-a interstitial relative to M&RS; base of 1m-cu&M separated from CU by about length of 2cu-a; hind wing with distal abscissa of CU present and complete; first abscissa of RS subequal to rs-m. Tarsal claw with proximal lobe quadrangular, with claw apex slightly overtaking the distal margin of lobe. Color. Head black except 0.8 distal of clypeus yellowish; antennae brownish with scape and pedicel ventrally whitish; mouthparts whitish, except apex of mandible brownish. Mesosoma orange with weak spot posteriorly of metapleuron and posterior carinae of propodeum brown. Metasoma blackish with anterior margin centrally orange in tergite I, tergites II-VI with anterior and anterolateral margins whitish. Fore leg orange, mid leg orange with coxa, trochanter and trochantellus whitish and tarsus brownish, hind leg whitish with coxa inner region, trochanter proximal, trochantellus distal, femur proximally and distally, tibia proximally and distally, first tarsal segment distally, and remaining tarsal segments entirely blackish brown. Wings hyaline, pterostigma brown. Ovipositor darkish brown, with posterior and anterior part whitish.
Male. (Fig. 6G). Similar to female in structure and coloration, but body about 5.0 mm; malar space 0.3 times as long as proximal mandibular width; lower face about 1.15 times as broad as high; lateral ocelli separated from compound eyes by about 0.75 times their own maximum diameter; fore wing length about 4.0 mm. Distribution. Brazil (Fig. 11). Biological note. Host unknown. Etymology. The specific name refers to the type locality of this species Serra dos Órgãos, Rio de Janeiro state, Brazil.
Remarks. Polysphincta organensis sp. nov. closely resembles P. bonita sp. nov. mainly by coloration: mesosoma entirely orange and propodeum brownish with some whitish in anterior part on tergites. It differs from P. bonita sp. nov. by having ovipositor >1.2 times as long as hind tibia and fore leg orange, mid leg orange with coxa, trochanter and trochantellus whitish and tarsus brownish, hind leg whitish with inner region of coxa, trochanter proximally, trochantellus distally, femur proximally and distally, tibia proximally and distally, first tarsal segment distally, and remaining tarsal segments entirely blackish brown (ovipositor 1.0 times as long as hind tibia and fore and mid leg orange, hind leg brownish, except coxa orange and a medium inner and outer region whitish in P. bonita sp. nov.).
Description. Female. Body [13.5] mm. Head. Clypeus weakly convex, posterior margin thin and flat centrally; malar space [0.4] times as long as proximal mandibular width; lower face about [1.0] times as broad as high, weakly convex centrally, polished, with fine sparse setiferous punctures; head in dorsal view with margin of gena flat behind eyes, and its margin about [0.5] times length of eye; ocelli moderately large, lateral ones separated from compound eyes by [0.85] times their own maximum diameter. Mesosoma. Pronotum without epomia; shelf-like projection, in dorsal view, more or less bilobed, broader than long, and, in lateral view, slender and with anterolateral corners weakly decurved; mesoscutum slender, in dorsal view, smooth and polished, with notauli weakly impressed anteriorly; scutellum convex, not laterally carinate; mesopleuron highly polished, virtually impunctate; epicnemial carina reaching almost level of lower corner of pronotum; epicnemium with vestigial vertical carina near lower corner of pronotum; metapleuron convex, smooth and polished, with few sparse, fine bristles evenly spaced, without discernible submeta- pleural carina. Propodeum mediodorsally smooth and polished, with longitudinal carinae present only posteriorly and laterally with scattered fine bristles. Fore wing length [10.0] mm; 1cu-a more or less interstitial relative to M&RS; base of 1m-cu&M separated from CU by more than length of 2cu-a; hind wing with distal abscissa of CU present and complete; first abscissa of RS subequal to rs-m. Tarsal claw with proximal lobe quadrangular, with claw apex slightly overtaking the distal margin of lobe. Distribution. Ecuador (Fig. 11). Biological note. Host unknown.
Etymology. The specific name (in apposition) refers to type locality of this species, Pichincha province, Ecuador.

Polysphincta shabui Gauld, 1991
Comments. Additional characters to the original description (♀) are as follows: body about [10.5-11.0] 10.5-14.0; head in dorsal view with margin of the gena flat behind the eyes, and its margin [0.5] 0.5-0.6 times length of eye; shelf-like projection, in dorsal view, developed anterolaterally in apex, the apex bilobed, and in lateral view, with anterolateral part in apex rounded and weakly decurved; mesoscutum robust, in dorsal view; tarsal claw with proximal lobe quadrangular, with claw apex slightly overtaking the distal margin of lobe.
Distribution. Costa Rica and Brazil (Fig. 12).  Diagnosis. Polysphincta sinearanea can be distinguished from other species of the P. dizardi species-group by the combination of the following characters: (1) epomia present, about 0.9 times length of proximal mandibular width (Fig. 9B); (2) malar space 0.5-0.6 times as long as proximal mandibular width (Fig. 9B); (3) fore wing vein 1cu-a more or less interstitial relative to M&RS (Fig. 9A); (4) mesosoma entirely orange (Fig. 9A); (5) fore wing very slightly yellowish hyaline (Fig. 9A); (6) hind leg orange with femur, tibia and tarsus brownish (Fig. 9A); (7) metasoma orange, with posterior margins of tergites II-IV narrowly black, posterior half of tergite V black, and tergites VI+ black (Fig. 9A, C); (8) ovipositor robust, 1.5 times as long as hind tibia. Comments. Additional characters to the original description (♀) are as follows: head in dorsal view with margin of the gena flat behind the eyes, and its margin [0.7] times length of eye; shelf-like projection, in dorsal view, weakly developed in the anterolateral part of apex, the apex more or less straight, not bilobed, and in lateral view, with anterolateral part in apex more or less rounded and not decurved; mesoscutum robust, in dorsal view; tarsal claw with proximal lobe quadrangular, with claw apex slightly overtaking the distal margin of lobe; ovipositor robust.
Distribution. Brazil (Fig. 12). (2) malar space 0.4-0.6 times as long as proximal mandibular width (Fig. 10B); (3) fore wing with vein 1cu-a postfurcal relative to M&RS (0.25-0.35 times its own length) or 1cu-a more or less interstitial relative to M&RS (Fig. 10A); (4) mesosoma orange, except posterior carinae of propodeum darkish brown (Fig. 10A); (5) fore wing hyaline (Fig. 10A); (6) hind leg whitish with proximal region of trochanter, longitudinal spot in subdistal region of inner and outer margin of femur, distal part of tibia and first tarsal segment distally and remaining tarsal segments brownish (Fig. 10A); (7) metasoma Description. Female. Body [7.75] 7.0-9.0 mm. Head. Clypeus weakly convex, posterior margin thin and straight centrally; malar space [0.4] 0.4-0.6 times as long as proximal mandibular width; lower face about [1.1] 0.9-1.1 times as broad as high, weakly convex centrally, polished, with fine sparse setiferous punctures; head in dorsal view with margin of gena very weakly convex behind eyes, and its margin [0.55] 0.4-0.6 times length of eye; ocelli moderately large, lateral ones separated from compound eyes by [0.9] 0.75-1.0 times their own maximum diameter. Mesosoma. Pronotum with epomia distinct, about [0.9] 0.9-1.3 times length of proximal mandibular width; shelf-like projection, in dorsal view, developed in anterolateral part of apex, apex bilobed, and in lateral view, with anterolateral part in apex rounded and weakly decurved; mesoscutum robust, in dorsal view, smooth and polished, with notauli weakly impressed anteriorly; scutellum convex, not laterally carinate; mesopleuron highly polished, virtually impunctate; epicnemial carina reaching almost level of lower corner of pronotum; epicnemium with vestigial vertical carina near lower corner of pronotum; metapleuron convex, smooth and polished, with sparse, fine bristles evenly spaced, without discernible submetapleural carina. Propodeum mediodorsally smooth and polished, with longitudinal carinae present only posteriorly and laterally with scattered fine bristles. Fore wing length [6.0] 5.0-7.0 mm; 1cu-a postfurcal relative to M&RS by [0.35] 0.25-0.35 times its own length; base of 1m-cu&M separated from CU by more than length of 2cu-a; hind wing with distal abscissa of CU present and complete but weakly pigmented; first abscissa of RS subequal to rs-m. Tarsal claw with proximal lobe quadrangular, with claw apex slightly overtaking distal margin of lobe. Metasoma. Tergite I [1.1] 1.1-1.7 times as long as posteriorly broad, dorsally with lateromedian longitudinal carinae only discernible at extreme anterior part; sternite I with weak swelling near hind rim, and with weak median longitudinal ridge anteriorly; tergite II about [1.3] 1.1-1.3 times as long as posteriorly broad, highly polished, at most with only fine setiferous punctures laterally; tergite III [1.0] 1.0-1.3 times as long as posteriorly broad, highly polished, at most with only fine setiferous punctures laterally; subgenital plate subquadrate. Ovipositor slightly slender, [1.7] 1.4-1.7 times as long as hind tibia, posteriorly evenly tapered to sharp point.
Color. Head black except lower face and clypeus brownish; antennae brown, except apex of scape and pedicel whitish; mouthparts white, except apex of mandible black. Mesosoma orange, except posterior carinae of propodeum darkish brown. Metasoma entirely darkish brown, with posterior margins of tergites II-V narrowly black. Legs whitish, fore leg with 0.7 distal of femur, tibia and tarsus weakly rufescent; mid leg with 0.3 distal of femur and tibia weakly rufescent, 0.2 distal of tarsomere I, distal half of tarsomere II, 0.8 distal tarsomere III and tarsomeres IV+ brownish; hind leg with proximal region of trochanter, longitudinal spot in subdistal region of inner and outer margin of femur, distal part of tibia and first tarsal segment distally and remaining tarsal segments brownish. Wings hyaline, pterostigma brown. Ovipositor brown, with anterior and posterior portions slightly whitish.
Male. Unknown. Variation. Some specimens with clypeus whitish and fore and mid legs with femur and tibia whitish, others have the fore leg entirely orange; the mid leg orange with tarsomeres brownish; the hind leg whitish, with inner margin of coxa, trochanter, trochantellus, proximal region and longitudinal spot in subdistal region of inner and outer margin of femur, proximal and distal part of tibia and all tarsus darkish brown.