Helictopleurus fungicola A, B habitus of male (A) and female (B) C, D lateral and anterior view of the aedeagus; basoventral lobes (BVLb) E endophallus: lamella copulatrix (LC) K, L details of the superior side of the membranous sac of the endophallus; Endophallic spines (EpS) O lateral view of the head: arrows indicating the absence of humps and tubercles. Helictopleurus peyrierasi stat. rest. F, G habitus of male (F) and female (G) H, I lateral and anterior view of the aedeagus J endophallus: superior (SpLC) and inferior (IpLC) part of the lamella copulatrix M, N details of the superior side of the membranous sac of the endophallus P lateral view of the head: arrows indicate the frontoclypeal hump and the postoccipital central tubercle.

 
 
  Part of: Rossini M, Montreuil O, Grebennikov V, Tarasov S (2021) Genome sequencing reveals extraordinary cephalic horns in the Madagascan dung beetle genus Helictopleurus (Coleoptera, Scarabaeinae): insight from a revision of fungicola species group. ZooKeys 1033: 63-79. https://doi.org/10.3897/zookeys.1033.63527