A–D Maximum likelihood nuclear gene trees; numbers adjacent to nodes indicate bootstrap support greater than 70%. Trees were rooted with Speleomaster lexi (not shown, see Suppl. material 2: Fig. S2–5). Blue text = C. leviathan, red text = C. behemoth. A Toll B F-box/LRR-repeat protein C phosphatase 2A protein D Neuromusculin E NeighborNet network generated from summary distances (ingroup nuclear data only) calculated in POFAD.