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  • Article title
  • Abstract
  • Keywords
  • Introduction
  • Materials and methods
    • Study area and sampling design
    • Data analysis
    • Species co-occurrence
  • Results
  • Discussion
    • The role of competition in carabid beetles: an open question
    • Insights from null model approaches
    • Patterns in the desert and typical steppes
    • Patterns in the meadow steppe
    • The effect of the zero-structure
    • Within-ecosystem analyses
    • What patterns of co-existence tell us about competition
  • Conclusions
  • Acknowledgements
  • References
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