Shells broken by Badister pictus A Cyclophorus herklotsi Martens, 1860 B Diplommatina sp. C Semisulcospira libertina (Gould, 1859) D, E Physa acuta F Gyraulus tokyoensis (Mori, 1938) G Hippeutis cantori H Polypylis hemisphaerula (Benson, 1842) I Pseudosuccinea columella (Say, 1817) J Opeas pyrgula K Zonitoides arboreus L Sinoennea iwakawa (Pilsbry, 1900) M Aegista aemula (Gude, 1900) N, O Euhadra idzumonis (Pilsbry & Gulock, 1900) P adult B. pictus A, F, G, H, K, M, N dorsal view B, C, D, J, L, O front view E, I back view. The soft bodies of these snails were eaten by B. pictus.

 
 
  Part of: Hayashi M, Sugiura S (2021) Shell-breaking predation on gastropods by Badister pictus (Coleoptera, Carabidae) with strikingly asymmetric mandibles. In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 815-830. https://doi.org/10.3897/zookeys.1044.62293