Attacking and feeding behavior of Badister pictus adults A adult biting the outer lip of Zonitoides arboreus (Say, 1816) B adult feeding on the soft body after opening the shell of Z. arboreus C adult breaking the shell of Opeas pyrgula Schmacker & Boettger, 1891 D adult biting the outer lip of Metalycaeus hirasei (Pilsbry, 1900) E adult feeding on the soft body from the broken shell of Hippeutis cantori (Benson, 1850) F adult biting the basal lip of the sinistral snail Physa acuta Draparnaud, 1805.

 
 
  Part of: Hayashi M, Sugiura S (2021) Shell-breaking predation on gastropods by Badister pictus (Coleoptera, Carabidae) with strikingly asymmetric mandibles. In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 815-830. https://doi.org/10.3897/zookeys.1044.62293