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  • Article title
  • Abstract
  • Keywords
  • Introduction
  • Materials and methods
    • Study species
    • Observation of beetle mandibles
    • Feeding experiments
    • Measurement of shell morphology
    • Data analysis
  • Results
    • Morphology of beetle mandibles
    • Feeding behavior
    • Effects of shell morphology
  • Discussion
  • Acknowledgements
  • References
  • Supplementary materials
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