Dorsal, lateral, frontal and declivital view of Coptoborus incomptus holotype, 1.7–1.9 mm (A–C, M), C. incultus holotype, 2.3 mm (D–F, N), C. inornatus paratype, 1.8 mm (G–I, O), C. janeway holotype, 2.0 mm (J–L, P). All photographs by SMS except G–I, O by T.H. Atkinson, copyright National Museum of Natural History, Smithsonian Institution, Washington, D.C., published by permission.