Dorsal, lateral, frontal and declivital view of Coptoborus osbornae holotype, 1.5–1.7 mm (A–C, M), C. panosus holotype, 2.4 mm (D–F, N), C. papillicauda holotype, 2.0 mm (G–I, O), C. paurus holotype, 1.7 mm (J–L, P). All photographs by SMS, except J–L, P copyright National Museum of Natural History, Smithsonian Institution, Washington, D.C., published by permission.