Tribe BLONDELIINI
Genus AFROLIXA Curran, 1939
AFROLIXA Curran, 1939: 4. Type species: Afrolixa macula Curran, 1939, by original designation.
macula Curran, 1939.—Afrotropical: Malawi, Mozambique, South Africa.
Afrolixa macula Curran, 1939: 4. Holotype male (SANC). Type locality: Mozambique, Maputo [as “Lourenco Marquis”].
Undescribed sp.: Côte d’Ivoire, Sudan, Uganda (BMNH, Crosskey 1984: 267).
Genus ANOMALOSTOMYIA Cerretti & Barraclough, 2007
ANOMALOSTOMYIA Cerretti & Barraclough, 2007: 102. Type species: Anomalostomyia namibica Cerretti & Barraclough, 2007, by original designation.
Note: Cerretti and Barraclough (2007: 104) considered Anomalostomyia as congeneric with Crosskey’s (1984: 289) “Undetermined genus”, which was based on a single male from Angola. That specimen, originally in BMNH, cannot be located (Cerretti and Barraclough 2007). Crosskey (1984: 289) treated the genus in Eryciini (and “apparently allied to the Erythrocera-group of genera”) but it has been provisionally placed in Blondeliini by Cerretti and Barraclough (2007) based on the species listed here from Namibia.
namibica Cerretti & Barraclough, 2007.—Afrotropical: Namibia.
Anomalostomyia namibica Cerretti & Barraclough, 2007: 103. Holotype male (NMNW). Type locality: Namibia, Brandberg Mountain, Sonusib Ravine, 1435m (21°04.546′S 14°36.958′E).
Genus BLONDELIA Robineau-Desvoidy, 1830
BLONDELIA Robineau-Desvoidy, 1830: 122. Type species: Blondelia nitida Robineau-Desvoidy, 1830 (= Tachina nigripes Fallén, 1810), by subsequent designation of Duponchel in d’Orbigny (1842: 609) (see Evenhuis and Thompson 1990: 233) [Palaearctic].
tibialis Mesnil, 1962.—Afrotropical: Burundi (new record, MZUR [PC]), D.R. Congo, South Africa.
Blondelia tibialis Mesnil, 1962: 753. Holotype male (IRSNB [not MRAC as published]). Type locality: D.R. Congo, Nord-Kivu, Kibati [ca. 1°36′S 29°16′E].
Genus CHARITELLA Mesnil, 1957
CHARITELLA Mesnil, 1957: 31. Type species: Charitella gracilis Mesnil, 1957, by monotypy [Oriental].
METADRINOMYIA Shima, 1980: 259. Type species: Metadrinomyia proclinata Shima, 1980, by original designation [Palaearctic]. Syn. n.
Note: Metadrinomyia Shima, 1980 was first recognized from the Afrotropical Region by Cerretti (2012: 325). It is here placed in synonymy with Charitella Mesnil, 1957.
nigrescens Mesnil, 1977.—Afrotropical: ?Madagascar, Malawi.
Charitella nigrescens Mesnil, 1977d: 325. Holotype female (CNC). Type locality: Malawi, Mt. Mulanje [as “Mt. Mlanje”].
whitmorei (Cerretti, 2012).—Afrotropical: Burundi, D.R. Congo. Comb. n.
Metadrinomyia whitmorei Cerretti, 2012: 325. Holotype male (MZUR). Type locality: Burundi, Kayanza [Province], Parc National de la Kibira, 2200m (2°53′25.9″S 29°27′25.4″E).
Note: The recently described Metadrinomyia whitmorei Cerretti, 2012 is moved here to Charitella Mesnil, 1957.
Undescribed sp. 1: Madagascar (TAU, examined by PC).
Undescribed sp. 2: Comoros (MNHN, examined by PC).
Genus COMPSILURA Bouché, 1834
COMPSILURA Bouché, 1834: 58. Type species: Tachina concinnata Meigen, 1824, by subsequent designation of Mik (1894: 52–53).
concinnata (Meigen, 1824).—Afrotropical: “widespread W. Afr. n.-e. Afr., E. Afr. & sthn Afr.” (Crosskey 1980b: 855), including Nigeria, South Africa. Palaearctic: C. Asia, Europe (all), Japan, M. East (all), N. Africa (Madeira), Pal. China, Russia (W. Russia, W. Siberia, E. Siberia), Transcaucasia. Oriental: India, Indonesia, Malaysia, Nepal, Orien. China, Philippines, Ryukyu Is., Taiwan, Thailand. Australasian: Australia, N. Australasian. Nearctic: introduced and widespread in northeast, also British Columbia to California.
Tachina concinnata Meigen, 1824: 412. Holotype female (NHMW, Herting 1972: 5). Type locality: not given (probably Germany, Hamburg [specimen from von Winthem]).
Phorocera selecta Curran, 1940: 6. Holotype male (SANC). Type locality: South Africa, KwaZulu-Natal.
solitaria (Curran, 1940).—Afrotropical: Zimbabwe.
Phorocera solitaria Curran, 1940: 6. Holotype male (AMNH). Type locality: Zimbabwe, Harare [as “Salisbury”].
Undescribed sp.: Madagascar (TAU, examined by PC).
Undetermined sp.: Burundi (MZUR, examined by PC).
Genus DOLICHOTARSINA Mesnil, 1977
DOLICHOTARSINA Mesnil, 1977d: 324. Type species: Dolichotarsina gracilis Mesnil, 1977, by original designation.
gracilis Mesnil, 1977.—Afrotropical: Madagascar.
Dolichotarsina gracilis Mesnil, 1977d: 325. Holotype female (MNHN). Type locality: Madagascar, Toamasina, Périnet [ca. 18°55′S 48°25′E].
Genus EOMEDINA Mesnil, 1960
EOMEDINA Mesnil, 1960b: 652. Type species: Eomedina grisescens Mesnil, 1960 (= Degeeria apicalis Curran, 1927), by original designation.
Note: See Cerretti and Wyatt (2006) for a diagnosis of Eomedina Mesnil, 1960 and a key to the two species.
apicalis (Curran, 1927).—Afrotropical: D.R. Congo, Kenya, Nigeria, Sierra Leone, Tanzania (new record, TAU [PC]), Uganda (new record, TAU [PC]).
Degeeria apicalis Curran, 1927c: 8. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
Eomedina grisescens Mesnil, 1960b: 651. Holotype female (BMNH). Type locality: D.R. Congo [as “Südafrika”, in error], Katanga, Bukama.
hamoyensis Cerretti & Wyatt, 2006.—Afrotropical: Namibia.
Eomedina hamoyensis Cerretti & Wyatt, 2006: 64. Holotype female (NMNW). Type locality: Namibia, Rundu District, Hamoye National Forest (18°12′S 19°43′E).
Genus EOPHYLLOPHILA Townsend, 1926
EOPHYLLOPHILA Townsend, 1926c: 19. Type species: Eophyllophila elegans Townsend, 1926, by original designation [Oriental].
africana Villeneuve, 1935.—Afrotropical: Angola, Burundi, Cameroon, Kenya, Malawi, Nigeria, Sierra Leone, Tanzania, Uganda.
Eophyllophila africana Villeneuve, 1935a: 136. Syntypes, 1 male and 1 female (not located). Type localities: Nigeria (Oshogbo) and Uganda (west Rwenzori Range [as “W. Ruwenzori”], 1800m).
Undescribed spp.: Kenya, Malawi, Uganda (all in TAU, examined by PC).
Genus ERYNNIOLA Mesnil, 1977
ERYNNIOLA Mesnil, 1977c: 179. Type species: Erynniola atricolor Mesnil, 1977, by original designation.
atricolor Mesnil, 1977.—Afrotropical: Madagascar.
Erynniola atricolor Mesnil, 1977c: 181. Holotype male (MNHN). Type locality: Madagascar, Toamasina, Périnet [ca. 18°55′S 48°25′E].
russipes Mesnil, 1977.—Afrotropical: Madagascar.
Erynniola russipes Mesnil, 1977c: 181. Holotype female (MNHN). Type locality: Madagascar, Toamasina, Périnet [ca. 18°55′S 48°25′E].
Genus FILISTEA Cerretti & O’Hara, gen. n.
FILISTEA Cerretti & O’Hara, gen. n. Type species: Viviania aureofasciata Curran, 1927, by present designation.
Note: This new genus is described in the New Taxa of Afrotropical Tachinidae section.
aureofasciata (Curran, 1927).—Afrotropical: Cameroon (new record, ZMHB [PC]), D.R. Congo, Nigeria, Uganda. Comb. n.
Viviania aureofasciata Curran, 1927c: 8. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
verbekei Cerretti & O’Hara, sp. n.—Afrotropical: Cameroon, D.R. Congo, Nigeria, Uganda.
Filistea verbekei Cerretti & O’Hara, sp. n. Holotype male (ZMHB). Type locality: Cameroon, Kumba [as “Johann-Albrechtshöhe”] (4°38′N 9°28′E).
Note: This new species is described in the New Taxa of Afrotropical Tachinidae section.
Genus ISTOCHETA Rondani, 1859
FALLENIA Meigen, 1838: 265 (junior homonym of Fallenia Meigen, 1820). Type species: Tachina longicornis Fallén, 1810, by subsequent designation of Coquillett (1910: 544) [Palaearctic].
ISTOCHETA Rondani, 1859: 157, 171. Type species: Istocheta frontosa Rondani, 1859 (as “Sp. Typ. novaFrontalis Mihi”, incorrect original spelling, see O’Hara et al. 2011: 101) (= Phorocera cinerea Macquart, 1850), by original designation [Palaearctic].
ISTOCHAETA Marschall, 1873: 334. Unjustified emendation of Istocheta Rondani, 1859 (see O’Hara et al. 2011: 101, 262).
HISTOCHAETA Brauer & Bergenstamm, 1891: 445 [also 1891: 141]. Unjustified emendation of Istocheta Rondani, 1859 (see O’Hara et al. 2011: 101).
PROSOPOFRONTINA Townsend, 1926c: 33. Type species: Prosopofrontina pulchra Townsend, 1926, by original designation [Oriental].
UROPHYLLINA Villeneuve, 1937c: 5 (as subgenus of Urophylloides Brauer & Bergenstamm, 1893). Type species: Urophylloides (Urophyllina) rufipes Villeneuve, 1937, by monotypy [Oriental].
ANUROPHYLLINA Mesnil, 1961: 693 (as subgenus of Urophyllina Villeneuve, 1937). Nomen nudum (proposed after 1930 without designation of type species from four included species) (see note below and Evenhuis et al. 2008: 6).
ANUROPHYLLINA Mesnil, 1977d: 322 (as subgenus of Urophyllina Villeneuve, 1937). Type species: Urophylloides bicolor Villeneuve, 1937, by original designation [Oriental].
Note: Herting (1984: 24) accepted Anurophyllina Mesnil, 1961 as an available name and designated Urophylloides bicolor Villeneuve, 1937 as type species. The availability of Anurophyllina Mesnil, 1961 vs. Anurophyllina Mesnil, 1977 was properly cited by O’Hara (1996: 121) and Evenhuis et al. (2008: 6) but O’Hara et al. (2009: 48) inadvertently followed Herting (1984).
cerina (Mesnil, 1977).—Afrotropical: Madagascar.
Urophyllina (Anurophyllina) cerina Mesnil, 1977d: 322. Holotype female (MNHN). Type locality: Madagascar, Antsiranana, Montagne d’Ambre [Parc National, ca. 12°36′S 49°8′E].
conifrons (Villeneuve, 1950).—Afrotropical: Uganda.
Degeeria conifrons Villeneuve, 1950: 2. Holotype male (IRSNB). Type locality: Uganda, Entebbe.
crucigera (Mesnil, 1977).—Afrotropical: Madagascar.
Urophyllina (Anurophyllina) crucigera Mesnil, 1977d: 322. Holotype male (MNHN). Type locality: Madagascar, Toamasina, Périnet [ca. 18°55′S 48°25′E].
flava (Curran, 1927).—Afrotropical: Kenya, Nigeria, Sierra Leone.
Viviania flava Curran, 1927f: 108. Holotype male (BMNH). Type locality: Sierra Leone, Njala [ca. 8°14′N 12°1'W].
Degeeria frontosa Villeneuve, 1950: 3. Holotype female (IRSNB). Type locality: Kenya, west side of Mt. Kenya, Ngare Rungai, 2000m.
Genus KINIATILIOPS Mesnil, 1955
KINIATILIOPS Mesnil, 1955: 365. Type species: Kiniatiliops elegans Mesnil, 1955 (= Lomatacantha nigrapex Mesnil, 1952), by monotypy.
bilineatus (Mesnil, 1952).—Afrotropical: D.R. Congo.
Lomatacantha bilineata Mesnil, 1952a: 11. Holotype female (not located). Type locality: D.R. Congo, Nord-Kivu, Kamatembe, 2100m [ca. 1°19′S 29°6′E].
nigrapex (Mesnil, 1952).—Afrotropical: D.R. Congo, Ethiopia, Kenya, Rwanda, Tanzania, Zambia.
Lomatacantha nigrapex Mesnil, 1952a: 13. Holotype male (MRAC). Type locality: D.R. Congo, Nord-Kivu, Rutshuru, 1285m.
Kiniatiliops elegans Mesnil, 1955: 365. Holotype male (MRAC). Type locality: Rwanda, Byumba [as “terr. Biumba”, a former territory], “Gatsibu” [probably Gatsibo, ca. 1°35′S 30°15′E], 1800m.
trispina Mesnil, 1959.—Afrotropical: Kenya.
Kiniatiliops trispina Mesnil, 1959: 14. Holotype female (SMNS). Type locality: Kenya, Lake Jipe.
Genus KINIATILLA Villeneuve, 1938
KINIATILLA Villeneuve, 1938c: 10. Type species: Kiniatilla tricincta Villeneuve, 1938, by original designation.
KINIATILIA. Incorrect subsequent spelling of Kiniatilla Villeneuve, 1938 (Mesnil 1952a: 14).
brevipalpis Mesnil, 1952.—Afrotropical: Burundi, D.R. Congo.
Kiniatilia brevipalpis Mesnil, 1952a: 14. Holotype male (MRAC). Type locality: D.R. Congo, Nord-Kivu, Beni to Lesse [Lesse is located northeast of Beni at ca. 0°45′N 29°46′E].
tricincta Villeneuve, 1938.—Afrotropical: Burundi, D.R. Congo, Rwanda, Uganda.
Kiniatilla tricincta Villeneuve, 1938c: 11. Lectotype female (IRSNB), by designation herein (see Lectotype Designations section). Type locality: D.R. Congo, Bas-Congo, Mayumbé [a highland area west of Rivière Congo], Kiniati.
Genus LATIGINELLA Villeneuve, 1936
LATIGINELLA Villeneuve, 1936a: 4. Type species: Latiginella rufogrisea Villeneuve, 1936, by monotypy.
handeni Verbeke, 1963.—Afrotropical: Malawi (new record, NMDA [PC]), Mozambique, Tanzania.
Latiginella handeni Verbeke, 1963: 176. Holotype female (MRAC). Type locality: Tanzania, Handeni, 350m.
rufogrisea Villeneuve, 1936.—Afrotropical: D.R. Congo, Kenya, Nigeria.
Latiginella rufogrisea Villeneuve, 1936a: 4. Holotype female (IRSNB). Type locality: Kenya, Ikutha.
Genus LINDNERIOLA Mesnil, 1959
LINDNERIOLA Mesnil, 1959: 17. Type species: Lindneriola paradoxa Mesnil, 1959, by monotypy.
paradoxa Mesnil, 1959.—Afrotropical: Tanzania, Uganda.
Lindneriola paradoxa Mesnil, 1959: 17. Holotype female (SMNS). Type locality: Tanzania, “Ngaruka” [probably Engaruka, ca. 3°0′S 35°58′E].
Undescribed sp. 1: South Africa (NMB, examined by PC).
Undescribed sp. 2: Tanzania (TAU, examined by PC).
Genus MAURITIODORIA Townsend, 1932
MAURITIODORIA Townsend, 1932: 52. Type species: Medoria spinicosta Thomson, 1869, by original designation.
GASTROLEPTINA Villeneuve, 1938c: 6. Type species: Gastroleptina discolor Villeneuve, 1938 (= Medoria spinicosta Thomson, 1869), by monotypy.
spinicosta (Thomson, 1869).—Afrotropical: Mauritius, Réunion.
Medoria spinicosta Thomson, 1869: 522. Lectotype male (NHRS), by fixation of Townsend (1932: 52) (examination of “Male Ht” from Mauritius in NHRS is regarded as a lectotype fixation). Type locality: Mauritius.
Clytia spinicosta Thomson, 1869: 523 (junior secondary homonym of Medoria spinicosta Thomson, 1869). Type(s), male (NHRS). Type locality: Mauritius.
Gastroleptina discolor Villeneuve, 1938c: 7. Syntypes, 1 male and 1 female (BMNH). Type locality: Mauritius.
Note: The relative priority of Medoria spinicosta Thomson, 1869 and Clytia spinicosta Thomson, 1869, when both are placed in the same genus, was established by Crosskey (1980b: 856), as the First Reviser (Article 24.2.2 of the Code, ICZN 1999). Townsend (1932: 52) was probably mistaken when he referred to the “male Pt” of Medoria spinicosta Thomson as bearing the label “Clytia spinicosta, Th”; this specimen is likely the name-bearing type of Clytia spinicosta Thomson, 1869.
Genus MEDINA Robineau-Desvoidy, 1830
MEDINA Robineau-Desvoidy, 1830: 138. Type species: Medina cylindrica Robineau-Desvoidy, 1830 (= Tachina collaris Fallén, 1820), by subsequent designation of Coquillett (1910: 565) [Palaearctic].
DEGEERIA Meigen, 1838: 249. Type species: Tachina collaris Fallén, 1820, by subsequent designation of Rondani (1856: 72) [Palaearctic].
carbonata Mesnil, 1968.—Afrotropical: Madagascar, South Africa, Tanzania.
Medina carbonata Mesnil, 1968b: 8. Holotype male (SMNS). Type locality: Tanzania, Makoa [probably near Moshi, ca. 3°21′S 37°19′E].
cinctella (Villeneuve, 1950).—Afrotropical: Malawi. Status revived.
Degeeria cinctella Villeneuve, 1950: 7. Holotype male (IRSNB). Type locality: Malawi, Mt. Mulanje [as “Mt. Mlanje”].
Note: Degeeria cinctella Villeneuve, 1950 was treated as a synonym of Medina lateralis (Villeneuve, 1950) by Verbeke (1964: 181) and Crosskey (1980b: 857) but is recognized here as a distinct species based on examination of the holotype by PC. The relative priority of Degeeria lateralis Villeneuve, 1950 and Degeeria cinctella Villeneuve, 1950, when the two are treated as synonyms, was established by Verbeke (1964: 181), as the First Reviser (Article 24.2.2 of the Code, ICZN 1999).
crocea (Villeneuve, 1950).—Afrotropical: Kenya, Malawi.
Degeeria crocea Villeneuve, 1950: 3. Lectotype male (IRSNB), by designation herein (see Lectotype Designations section). Type locality: Malawi, Mt. Mulanje [as “Mt. Mlanje”]).
decellei Verbeke, 1964.—Afrotropical: Côte d’Ivoire.
Medina decellei Verbeke, 1964: 169. Holotype male (MRAC). Type locality: Côte d’Ivoire, Parc du Banco [as “Réserve du Banco”; near Abidjan].
denticulata (Villeneuve, 1950).—Afrotropical: Madagascar, Nigeria.
Degeeria denticulata Villeneuve, 1950: 6. Holotype female (IRSNB). Type locality: Nigeria, Ilesha.
egregia (Villeneuve, 1950).—Afrotropical: Nigeria, Zambia, Zimbabwe.
Degeeria egregia Villeneuve, 1950: 4. Holotype male (IRSNB). Type locality: Nigeria, Oshogbo.
lateralis (Villeneuve, 1950).—Afrotropical: Burundi, D.R. Congo (new record, IRSNB [PC]), Rwanda, South Africa, Tanzania.
Degeeria lateralis Villeneuve, 1950: 7. Holotype male (IRSNB). Type locality: South Africa, Western Cape, Cape of Good Hope [as “Cap”].
mira Mesnil, 1977.—Afrotropical: Madagascar.
Medina mira Mesnil, 1977c: 185. Holotype male (MNHN). Type locality: Madagascar, Mahajanga, Ambato Boeni.
nigra Mesnil, 1968.—Afrotropical: Angola, Madagascar, South Africa.
Medina nigra Mesnil, 1968b: 8. Holotype male (SMNS). Type locality: South Africa, Western Cape, Cape Town.
pectinifera Mesnil, 1977.—Afrotropical: Madagascar.
Medina pectinifera Mesnil, 1977c: 187. Holotype female (MNHN). Type locality: Madagascar, Antsiranana, Montagne d’Ambre [Parc National, ca. 12°36′S 49°8′E].
rubricosa (Villeneuve, 1913).—Afrotropical: Nigeria.
Lydella rubricosa Villeneuve, 1913c: 30. Holotype female (BMNH). Type locality: Nigeria, Oshogbo.
Note: Villeneuve’s (1913c: 30–31) description of a single female of Lydella rubricosa from Nigeria was followed by a brief description of a male from Benin (as “Dahomey”). It is not clear whether this male was thought to be conspecific with L. rubricosa and hence part of the type series of this nominal species. We have inferred that the male was not positively associated with the female and is therefore not a syntype of L. rubricosa, and have followed Crosskey (1980b: 857) in excluding Benin from the distribution of M. rubricosa (Villeneuve).
semirufa (Villeneuve, 1950).—Afrotropical: Kenya, Malawi.
Degeeria semirufa Villeneuve, 1950: 6. Lectotype female (IRSNB), by designation herein (see Lectotype Designations section). Type locality: Malawi, Mt. Mulanje [as “Mt. Mlanje”].
setosella (Villeneuve, 1950).—Afrotropical: Burundi (new record, IRSNB [PC]), Cameroon, D.R. Congo (new record, IRSNB [PC]), Uganda.
Degeeria setosella Villeneuve, 1950: 5. Holotype male (IRSNB). Type locality: northwest Cameroon, Dschang [as “Dchang”] Plateau.
sopha Mesnil, 1977.—Afrotropical: Madagascar.
Medina sopha Mesnil, 1977c: 184. Holotype male (MNHN). Type locality: Madagascar, Toamasina, Périnet, 1000m [ca. 18°55′S 48°25′E].
spinulifera Mesnil, 1968.—Afrotropical: Tanzania.
Medina spinulifera Mesnil, 1968b: 9. Holotype female (SMNS). Type locality: Tanzania, Makoa [probably near Moshi, ca. 3°21′S 37°19′E].
succuba Mesnil, 1977.—Afrotropical: Madagascar.
Medina succuba Mesnil, 1977c: 186. Holotype male (MNHN). Type locality: Madagascar, Toamasina, Moramanga.
vidua Mesnil, 1977.—Afrotropical: Madagascar.
Medina vidua Mesnil, 1977c: 187. Holotype female (MNHN). Type locality: Madagascar, Toamasina, Périnet [ca. 18°55′S 48°25′E].
Possibly undescribed sp.: Nigeria (Crosskey 1984: 265).
Genus MEIGENIA Robineau-Desvoidy, 1830
MEIGENIA Robineau-Desvoidy, 1830: 198. Type species: Meigenia cylindrica Robineau-Desvoidy, 1830, by subsequent designation of Desmarest in d’Orbigny (1849a: 318, as “T. [Tachina] cylindrica”) (see Evenhuis and Thompson 1990: 237) [Palaearctic].
Note: Meigenia cylindrica Robineau-Desvoidy, 1830 is accepted as the type species of Meigenia Robineau-Desvoidy, 1830, following Evenhuis and Thompson (1990: 237). This name was treated as a nomen dubium under Meigenia by Herting and Dely-Draskovits (1993: 147). Despite this treatment of the type species of Meigenia as a nomen dubium, the concept of Meigenia is well-established and no useful purpose would be served by calling it into question over the dubious identity of M. cylindrica.
Undetermined sp.: Yemen (Zeegers 2007: 388).
Genus MEDINOSPILA Mesnil, 1977
MEDINOSPILA Mesnil, 1977d: 322. Type species: Medinospila nigella Mesnil, 1977, by original designation.
nigella Mesnil, 1977.—Afrotropical: Madagascar.
Medinospila nigella Mesnil, 1977d: 323. Holotype male (MNHN). Type locality: Madagascar, Toamasina, Périnet [ca. 18°55′S 48°25′E].
Genus PARARONDANIA Villeneuve, 1916
PARARONDANIA Villeneuve, 1916c: 498. Type species: Pararondania multipunctata Villeneuve, 1916, by monotypy.
multipunctata Villeneuve, 1916.—Afrotropical: South Africa.
Pararondania multipunctata Villeneuve, 1916c: 498. Holotype female (CNC [not SAMC as published]). Type locality: South Africa, “Cape Colony” (“S.W. Distr. Cape Col.” according to label data, Cooper and O’Hara 1996: 58; possibly referring to present-day Western Cape, Cape of Good Hope).
Genus PARATRIXA Brauer & Bergenstamm, 1891
PARATRIXA Brauer & Bergenstamm, 1891: 357 [also 1891: 53]. Type species: Paratrixa polonica Brauer & Bergenstamm, 1891, by monotypy [Palaearctic]. New record.
Note: Mesnil (1952a) described the two Afrotropical species below in Paratrixa Brauer & Bergenstamm, 1891. Crosskey (1980b: 857) did not recognize Paratrixa and placed these two species in Medina Robineau-Desvoidy, 1830. Paratrixa is treated as a genus in the Palaearctic Region (e.g., Herting and Dely-Draskovits 1993: 153, Cerretti 2010: 128) and is reinstated here as an Afrotropical genus with these same two species.
aethiopica Mesnil, 1952.—Afrotropical: D.R. Congo, Rwanda, South Africa. Comb. revived.
Paratrixa aethiopica Mesnil, 1952a: 10. Holotype female (not located). Type locality: Rwanda, Ruhengeri [1°30′S 29°38′E], “sources Kirii” [not located], 1800–1825m.
stammeri Mesnil, 1952.—Afrotropical: D.R. Congo, South Africa (new record, IRSNB [PC]). Comb. revived.
Paratrixa stammeri Mesnil, 1952a: 9. Holotype male (not located). Type locality: D.R. Congo, Nord-Kivu, Rutshuru, 1285m.
Genus PELASHYRIA Villeneuve, 1935
PELASHYRIA Villeneuve, 1935a: 138. Type species: Pelashyria grisescens Villeneuve, 1935, by monotypy.
grisescens Villeneuve, 1935.—Afrotropical: D.R. Congo.
Pelashyria grisescens Villeneuve, 1935a: 139. Syntypes, 1 male and 1 female (IRSNB). Type locality: D.R. Congo, Nord-Kivu, Mukule, 1800m [ca. 1°20′S 29°15′E].
Genus PRODEGEERIA Brauer & Bergenstamm, 1894
PRODEGEERIA Brauer & Bergenstamm, 1894: 617 [also 1895: 81]. Type species: Prodegeeria javana Brauer & Bergenstamm, 1894, by monotypy [Oriental].
MYXHYPOSTENA Villeneuve, 1939: 6. Type species: Myxhypostena consobrina Villeneuve, 1939, by original designation.
Note: Villeneuve (1939: 6) wrote about his new genus Myxhypostena: “le scutellum du type à 4 soies marginales”. This statement is accepted as a type species designation for Myxhypostena of the single included species, Myxhypostena consobrina Villeneuve.
consobrina (Villeneuve, 1939).—Afrotropical: D.R. Congo, Ghana, Nigeria.
Myxhypostena consobrina Villeneuve, 1939: 6. Syntypes, 1 male and 1 female (IRSNB). Type localities: D.R. Congo (“Agangula” [not located]) and Nigeria (Oshogbo).
straeleni Mesnil, 1952.—Afrotropical: D.R. Congo, Uganda.
Prodegeeria straeleni Mesnil, 1952a: 14. Holotype male (IRSNB). Type locality: D.R. Congo, Équateur, Eala.
Genus PROSUCCINGULUM Mesnil, 1959
PROSUCCINGULUM Mesnil, 1959: 16. Type species: Prosuccingulum aberrans Mesnil, 1959, by monotypy.
aberrans Mesnil, 1959.—Afrotropical: Tanzania.
Prosuccingulum aberrans Mesnil, 1959: 16. Holotype female (SMNS). Type locality: Tanzania, west side of Mt. Kibo [one of the three peaks of Mt. Kilimanjaro], 2800m.
Undescribed sp.: Malawi (NMB, examined by PC).
Genus RIOTERIA Herting, 1973
RIOTERIA Herting, 1973: 3. Type species: Rioteria submacula Herting, 1973, by monotypy [Palaearctic].
flava Zeegers, 2007.—Afrotropical: Yemen.
Rioteria flava Zeegers, 2007: 395. Holotype male (RMNH). Type locality: Yemen, 12km northwest of Manākhah (15°04′19″N 43°44′27″E).
rufitibia (Mesnil, 1959).—Afrotropical: Nigeria, Tanzania.
Tachinophytopsis rufitibia Mesnil, 1959: 14. Holotype male (SMNS). Type locality: Tanzania, “Ngaruka” [probably Engaruka, ca. 3°0′S 35°58′E].
Undescribed sp. 1: South Africa (NMB, examined by PC).
Undescribed sp. 2: Burkina (MZUR, examined by PC).
Genus TRIGONOSPILA Pokorny, 1886
TRIGONOSPILA Pokorny, 1886: 191. Type species: Trigonospila picta Pokorny, 1886 (= Tachina ludio Zetterstedt, 1849), by monotypy [Palaearctic].
SUCCINGULUM Pandellé, 1894: 52. Type species: Succingulum transvittatum Pandellé, 1896, by subsequent monotypy of Pandellé (1896: 148) [Palaearctic].
bimaculata (Villeneuve, 1935).—Afrotropical: Ghana, Malawi, Mozambique, Nigeria, Sudan, Uganda.
Succingulum bimaculatum Villeneuve, 1935a: 142. Holotype female (IRSNB). Type locality: Malawi.
Note: Villeneuve (1935a: 142) cited a second female of Succingulum bimaculatum seen by W.S. Patton but it was not examined by Villeneuve (as evidenced from his statement, “La tarière est exserte sur l’unique ♀ que j’ai vue”) and hence is not a syntype.
exigua (Villeneuve, 1935).—Afrotropical: South Africa. Status revived.
Succingulum exiguum Villeneuve, 1935a: 142. Holotype male (IRSNB). Type locality: South Africa.
Note: Succingulum exiguum Villeneuve, 1935 was treated as a synonym of Trigonospila mista (Villeneuve, 1913) by Crosskey (1980b: 858) but is recognized here as a distinct species based on examination of the holotype by PC.
integra (Villeneuve, 1935).—Afrotropical: “Afrique”. Oriental: India, Myanmar.
Succingulum integrum Villeneuve, 1935a: 142. Holotype male (possibly lost, Crosskey 1976: 218). Type locality: Africa [as “Afrique (région?)”].
mista (Villeneuve, 1913).—Afrotropical: Angola, D.R. Congo, Kenya, Malawi, ?South Africa, Tanzania, Uganda.
Succingulum mista Villeneuve, 1913c: 39. Holotype female (IRSNB). Type locality: D.R. Congo, Katanga, Sankisia.
prasius Mesnil, 1977.
prasiusprasius Mesnil, 1977.—Afrotropical: Madagascar.
Trigonospila prasius prasius Mesnil, 1977c: 181, 183. Holotype male (MNHN). Type locality: Madagascar, Toamasina, Périnet [ca. 18°55′S 48°25′E].
prasius trifida Mesnil, 1977.—Afrotropical: Madagascar.
Trigonospila prasius trifidus Mesnil, 1977c: 183. Holotype male (MNHN). Type locality: Madagascar, Antananarivo, Antananarivo [as “Tananarive”].
Unplaced species of Blondeliini
triquetra Macquart, 1844.—Afrotropical: Réunion.
Dexia triquetra Macquart, 1844: 86 [also 1844: 243]. Lectotype male (MNHN), by fixation of Crosskey (1971: 267) (examination of “Holotype ♂” from Réunion in MNHN is regarded as a lectotype fixation). Type locality: Réunion [as “Bourbon”].
Tribe ERYCIINI
Genus AFROPHYLAX Cerretti & O’Hara, gen. n.
AFROPHYLAX Cerretti & O’Hara, gen. n. Type species: Sturmia aureiventris Villeneuve, 1910, by present designation.
Note: This new genus is described in the New Taxa of Afrotropical Tachinidae section.
aureiventris (Villeneuve, 1910).—Afrotropical: Cameroon (new record, ZMHB [PC]), D.R. Congo, Nigeria, Sierra Leone, Tanzania, Uganda. Comb. n.
Sturmia aureiventris Villeneuve, 1910a: 252. Holotype male (MRAC). Type locality: D.R. Congo (as “Congo”, p. 249).
Note: Villeneuve (1910a) described four species from “Congo”. Curran (1927f: 122) treated one of them (Sturmia aureiventris Villeneuve, 1910) as described from D.R. Congo (as “Belgian Congo”), and used “Belgian Congo” and “Congo” interchangeably in this work and some others. We think it likely that Villeneuve (1910a), like Curran, used “Congo” in the sense of present-day D.R. Congo. However, Crosskey (1980b) interpreted Villeneuve’s Congo as the present-day country of Congo. Crosskey (1980b: 867, 1984: 277) treated Sturmia aureiventris Villeneuve as an unplaced species in the “Carceliini”.
Genus ANTISTASEA Bischof, 1904
ANTISTASEA Bischof, 1904: 82. Type species: Antistasea fimbriata Bischof, 1904, by monotypy.
fimbriata Bischof, 1904.—Afrotropical: Kenya (new record, TAU [PC]), South Africa, Zimbabwe.
Antistasea fimbriata Bischof, 1904: 83. Lectotype male (NHMW), by fixation of Townsend (1941: 235) (mention of “Ht male” from Algoa Bay in NHMW is regarded as a lectotype fixation). Type locality: South Africa, Eastern Cape, Algoa Bay.
Podomyia discalis Curran, 1939: 2. Holotype male (AMNH). Type locality: Zimbabwe, Harare [as “Salisbury”]. Syn. n.
Note: Crosskey (1984: 289) commented that Podomyia discalis Curran, 1939 is “almost certainly synonymous” with Antistasea fimbriata Bischof, 1904. We confirm from examination of the name-bearing types that these names are synonyms.
mutans Mesnil, 1970.—Afrotropical: Botswana, South Africa.
Antistasea mutans Mesnil, 1970b: 106. Holotype male (CNC). Type locality: South Africa, KwaZulu-Natal, Mfongosi [ca. 28°43′S 30°49′E].
Genus APLOMYA Robineau-Desvoidy, 1830
APLOMYA Robineau-Desvoidy, 1830: 184. Type species: Aplomya zonata Robineau-Desvoidy, 1830 (= Tachina confinis Fallén, 1820), by subsequent designation of Robineau-Desvoidy (1863a: 459, 460) (as confinis, with zonata in synonymy) [Palaearctic].
APLOMYIA Agassiz, 1846a: 3. Unjustified emendation of Aplomya Robineau-Desvoidy, 1830 (see Evenhuis et al. 2010: 39).
HAPLOMYIA Agassiz, 1846b: 172. Unjustified emendation of Aplomya Robineau-Desvoidy, 1830 (see Evenhuis et al. 2010: 39).
PROZENILLIA Villeneuve, 1916c: 487. Type species: Prozenillia distans Villeneuve, 1916, by monotypy.
WIEDEMANNIOMYIA Townsend, 1933: 469. Type species: Tachina metallica Wiedemann, 1824, by original designation.
APLOMYIELLA Mesnil, 1939d: 31. Type species: Tricholyga impexa Villeneuve, 1916 (= Tachina metallica Wiedemann, 1824), by original designation.
ATRICHOLYGA Villeneuve, 1939: 9. Type species: Tricholyga impexa Villeneuve, 1916 (= Tachina metallica Wiedemann, 1824), by monotypy.
confinis (Fallén, 1820).—Afrotropical: ?Malawi, Yemen. Palaearctic: C. Asia, Europe (all), Japan, M. East (all), Mongolia, N. Africa (Canary Is., Madeira), Pal. China, Russia (W. Russia, W. Siberia, E. Siberia, S. Far East), Transcaucasia. Oriental: Orien. China.
Tachina confinis Fallén, 1820: 32. Syntypes, males and females (NHRS and/or MZLU). Type locality: Sweden, Gotland.
Note: Tachina confinis Fallén, 1820 was recorded from Malawi by Villeneuve (1913c: 32) but not by Crosskey (1980b: 876). The presence of this species in Malawi needs confirmation.
distans (Villeneuve, 1916).—Afrotropical: Nigeria, South Africa, Sudan, Uganda.
Prozenillia distans Villeneuve, 1916c: 488. Lectotype male (SAMC, not located by JEOH), by fixation of Townsend (1940: 311) (mention of “Ht male” from Durban in SAMC is regarded as a lectotype fixation, if type can be found in SAMC). Type locality: South Africa, KwaZulu-Natal, Durban.
latimana Villeneuve, 1934.—Afrotropical: D.R. Congo, Kenya, Uganda.
Aplomyia latimana Villeneuve, 1934c: 409. Holotype female (CNC). Type locality: Uganda, Rwenzori Range [as “Ruwenzori”], 1800m.
lycaena (Curran, 1927).—Afrotropical: Ethiopia, Senegal, South Africa.
Zenillia lycaena Curran, 1927d: 333. Holotype male (SANC). Type locality: South Africa, Free State, Bloemfontein.
metallica (Wiedemann, 1824).—Afrotropical: “W. Afr. to n.-e. Afr., E. Afr. & sthn Afr.” (Crosskey 1980b: 876), including D.R. Congo, Mozambique, South Africa, Sudan, U.A. Emirates, Yemen. Palaearctic: Japan, M. East (Israel), Pal. China. Oriental: India, Indonesia, Orien. China, Ryukyu Is., Taiwan. Australasian: N. Australasian.
Tachina metallica Wiedemann, 1824: 46. Lectotype male (ZMUC), by fixation of Townsend (1933: 470) (examination of “Male holotype” from East Indies in ZMUC is regarded as a lectotype fixation). Type locality: “India orient.” (i.e., “East Indies”).
Tachina nigriventris Wiedemann, 1824: 43. Lectotype male (ZMUC), by fixation of Townsend (1933: 470) (examination of “male holotype” from East Indies in ZMUC is regarded as a lectotype fixation). Type locality: “India orient.” (i.e., “East Indies”).
Tachina notata Wiedemann, 1830: 653. Type(s), male (SMF or lost). Type locality: Nubia region [as “Nubien”, a region in southern Egypt and northern Sudan, recorded here as Sudan following Crosskey 1980b: 876].
Tachina socia Wiedemann, 1830: 654. Type(s), female (SMF or lost). Type locality: not given (likely Nubia region).
Phorocera eucalypta Loew, 1852: 659 [also 1862: 19, full description]. Type(s), unspecified sex (1 male in ZMHB). Type locality: Mozambique (Tete [as “Tette”] according to Loew 1862: 20).
Parexorista laeviventris van der Wulp, 1893: 173. Lectotype male (RMNH), by designation of Crosskey (1966a: 674–675) (see also Crosskey 1969: 105). Type locality: Indonesia, Jawa.
Tricholyga impexa Villeneuve, 1916c: 494. Syntypes, 2 males (1 male in NHMW). Type localities: D.R. Congo [as “Congo”, but received from Bequaert and presumably collected from D.R. Congo] and South Africa (Eastern Cape, Uitenhage).
Note: The relative priority of Tachina metallica Wiedemann, 1824 and Tachina nigriventris Wiedemann, 1824, when the two are treated as synonyms, was established by Townsend (1933: 470), as the First Reviser (Article 24.2.2 of the Code, ICZN 1999). Tachina notata Wiedemann, 1830 and Tachina socia Wiedemann, 1830 were synonymized with T. metallica by Crosskey (1980b: 876); their relative priority has not been established by a First Reviser and such action is unnecessary while they are invalid names.
The male syntype of Tricholyga impexa Villeneuve, 1916 in NHMW was collected from Uitenhage, South Africa, on 15 November 1896 and not on 15 December 1896 as given by Villeneuve (1916c: 494) (examined by JEOH).
poultoni (Villeneuve, 1922).—Afrotropical: Kenya, Nigeria, South Africa.
Exorista poultoni Villeneuve, 1922a: 518. Holotype male (not located). Type locality: Nigeria, near Ibadan, Moor Plantation.
seyrigi Mesnil, 1954.—Afrotropical: Madagascar.
Aplomyia (Aplomyiella) seyrigi Mesnil, 1954: 330. Holotype male (MNHN). Type locality: Madagascar, Toliara, Bekily.
versicolor (Curran, 1927).—Afrotropical: South Africa, Uganda.
Zenillia versicolor Curran, 1927d: 334. Holotype male (SANC). Type locality: South Africa, Eastern Cape, East London.
Genus CADURCIELLA Villeneuve, 1927
CADURCIELLA Villeneuve, 1927: 120. Type species: Cadurciella rufipalpis Villeneuve, 1927, by monotypy.
rufipalpis Villeneuve, 1927.—Afrotropical: Namibia, South Africa, Zimbabwe. Palaearctic: M. East (Israel).
Cadurciella rufipalpis Villeneuve, 1927: 120. Lectotype male (not located), by fixation of Townsend (1941: 248) (mention of “Ht male” from Salisbury in Rambouillet [Villeneuve’s personal collection, since dispersed] is regarded as a lectotype fixation for the single male in the type series from this locality). Type locality: Zimbabwe, Harare [as “Salisbury”].
uniseta (Curran, 1933).—Afrotropical: South Africa, Zimbabwe.
Zenillia uniseta Curran, 1933: 166. Holotype male (BMNH). Type locality: Zimbabwe.
Undetermined sp.: U.A. Emirates, as “cf. Cadurciella spec.” (Zeegers 2010: 681).
Genus CARCELIA Robineau-Desvoidy, 1830
Subgenus CARCELIA Robineau-Desvoidy, 1830
CARCELIA Robineau-Desvoidy, 1830: 176. Type species: Carcelia bombylans Robineau-Desvoidy, 1830, by subsequent designation of Coquillett (1910: 518) (see Evenhuis et al. 2010: 52) [Palaearctic].
CARCELLIA. Incorrect subsequent spelling of Carcelia Robineau-Desvoidy, 1830 (Rondani 1859: 103, Stackelberg 1943: 163) (see O’Hara et al. 2011: 46).
nudioculata Villeneuve, 1938.—Afrotropical: D.R. Congo, Rwanda, Uganda.
Carcelia nudioculata Villeneuve, 1938c: 4. Holotype male (not located). Type locality: D.R. Congo, Maniema, Lubutu.
Subgenus CARCELITA Mesnil, 1975
CARCELITA Mesnil, 1975a: 1384. Type species: Carcelia peraequalis Mesnil, 1950, by monotypy.
CARICELIA Mesnil, 1975a: 1384. Nomen nudum (proposed after 1930 without designation of type species; no included species).
CARICELIA Mesnil, 1975b: 1388. Type species: Carcelia obliterata Mesnil, 1950, by original designation.
Note: See O’Hara (1996: 122) for an explanation of the nomenclatural history of Caricelia Mesnil and Carcelita Mesnil.
abrelicta Mesnil, 1950.—Afrotropical: Burundi, D.R. Congo, South Africa, Tanzania, Uganda.
Carcelia abrelicta Mesnil, 1950b: 16. Syntypes, males and females (1 female in CNC). Type localities: D.R. Congo and South Africa (Western Cape, Cape Town).
aequalis Villeneuve, 1939.—Afrotropical: Nigeria, South Africa, Tanzania, Zimbabwe.
Carcelia aequalis Villeneuve, 1939: 1. Syntypes, males (“plusieurs individus”) (1 male in CNC, 1 male in SAMC). Type locality: South Africa, KwaZulu-Natal.
Note: One male of Carcelia aequalis Villeneuve, 1939 in IRSNB from “Stella B” [former Stella Bush near Durban] (examined by PC) with a Villeneuve determination label is likely an unmarked syntype.
angulicornis Villeneuve, 1916.—Afrotropical: Ghana (new record, CNC), Malawi, Nigeria, Sierra Leone, South Africa.
Carcelia angulicornis Villeneuve, 1916c: 481. Syntypes, males and females (BMNH, CNC). Type localities: Malawi (Mulanje [as “Mlange”]), Nigeria (Oshogbo), and South Africa.
argyriceps (Curran, 1927).—Afrotropical: Uganda.
Zenillia argyriceps Curran, 1927d: 328. Holotype male (BMNH). Type locality: Uganda, [Kanungu District in southwestern Uganda], Kinkizi County, “Kizazi” [not located].
Zenillia hargreavesi Curran, 1928a: 238. Holotype male (BMNH). Type locality: Uganda, Kampala.
atricans Mesnil, 1955.—Afrotropical: Burundi (new record, CNC, MZUR [PC]), ?Cape Verde, Kenya (new record, CNC), Rwanda, Tanzania.
Carcelia atricans Mesnil, 1955: 362. Holotype male (MRAC). Type locality: Rwanda, eastern foothills of Volcan Muhabura [as “Muhavura”], 2100m [ca. 1°23′S 29°44′E].
bigoti (Jaennicke, 1867).—Afrotropical: Ethiopia.
Exorista bigoti Jaennicke, 1867: 384 [also 1868: 76]. Type(s), female (SMF). Type locality: Ethiopia, “Simen” (probably the Simien Mountains area).
forcipata Mesnil, 1977.—Afrotropical: Madagascar.
Carcelia (Carcelita) forcipata Mesnil, 1977b: 178. Holotype male (MNHN). Type locality: Madagascar, Antananarivo, Ampefy.
inusta Mesnil, 1950.—Afrotropical: Malawi (new record, CNC), South Africa.
Carcelia inusta Mesnil, 1950b: 11. Syntypes, males and females (1 male in CNC). Type locality: South Africa, KwaZulu-Natal, “Stella” [former Stella Bush near Durban].
keiseri Mesnil, 1977.—Afrotropical: Madagascar.
Carcelia (Carcelita) keiseri Mesnil, 1977b: 176. Holotype male (MNHN). Type locality: Madagascar, Toamasina, Périnet, 1000m [ca. 18°55′S 48°25′E].
lindneri Mesnil, 1959.—Afrotropical: South Africa (new record, CNC), Tanzania.
Carcelia lindneri Mesnil, 1959: 2. Holotype male (SMNS). Type locality: Tanzania, Msingi [ca. 4°20′S 34°34′E].
lucidula Villeneuve, 1941.—Afrotropical: C.A. Republic (new record, CNC), D.R. Congo.
Carcelia lucidula Villeneuve, 1941b: 125. Syntypes, 2 males and 1 female (2 males in MRAC, 1 female in CNC). Type locality: D.R. Congo, Orientale, Uele, Dembia.
normula (Curran, 1927).—Afrotropical: D.R. Congo, Ghana, Nigeria, Tanzania, Uganda.
Zenillia normula Curran, 1927d: 329. Holotype female (BMNH). Type locality: Uganda, “Rosaka” [not located].
oblectanea Mesnil, 1950.—Afrotropical: D.R. Congo, Kenya, South Africa (new record, CNC).
Carcelia oblectanea Mesnil, 1950b: 15. Syntypes, males and females (1 female in CNC). Type locality: D.R. Congo.
oblimata Mesnil, 1950.—Afrotropical: South Africa.
Carcelia oblimata Mesnil, 1950b: 14. Syntypes, males and females (1 female in CNC). Type locality: South Africa, Western Cape, Cape Town.
obliterata Mesnil, 1950.—Afrotropical: Rwanda, South Africa.
Carcelia obliterata Mesnil, 1950b: 13. Lectotype female (CNC), by fixation of O’Hara (1996: 150). Type locality: South Africa (“Kransp.” according to label data, Cooper and O’Hara 1996: 21).
Note: O’Hara (1996: 150) accepted the specimen labelled as “TYPE” in CNC as the holotype of Carcelia obliterata Mesnil, 1950 under the assumption that the species was likely described from a single specimen. This assumption is contrary to Recommendation 73F, “Avoidance of assumption of holotype”, of the current Code (ICZN 1999). O’Hara’s (1996: 150) treatment of the “TYPE” in CNC as the holotype of C. obliterata is regarded as a lectotype fixation.
oculata (Villeneuve, 1910).—Afrotropical: D.R. Congo.
Exorista oculata Villeneuve, 1910a: 251. Lectotype male (IRSNB), by designation herein (see Lectotype Designations section). Type locality: D.R. Congo (as “Congo”, p. 249).
occulata. Incorrect subsequent spelling of oculata Villeneuve, 1910 (Curran 1927d: 335).
Note: Villeneuve (1910a) described four species from “Congo”. Curran (1927f: 122) treated one of them (Sturmia aureiventris Villeneuve, 1910) as described from D.R. Congo (as “Belgian Congo”), and used “Belgian Congo” and “Congo” interchangeably in this work and some others. We think it likely that Villeneuve (1910a), like Curran, used “Congo” in the sense of present-day D.R. Congo. However, Crosskey (1980b) interpreted Villeneuve’s Congo as the present-day country of Congo.
orbitalis (Curran, 1927).—Afrotropical: South Africa, Zimbabwe.
Zenillia orbitalis Curran, 1927d: 330. Holotype male (SANC). Type locality: South Africa, Gauteng, Pretoria.
patellata Mesnil, 1977.—Afrotropical: Madagascar.
Carcelia (Carcelita) patellata Mesnil, 1977b: 177. Holotype female (MNHN). Type locality: Madagascar, Antsiranana, Montagne d’Ambre [Parc National, ca. 12°36′S 49°8′E].
pellex Mesnil, 1950.—Afrotropical: Kenya, South Africa, Uganda.
Carcelia pellex Mesnil, 1950b: 13. Type(s), unspecified sex (not located). Type locality: South Africa.
peraequalis Mesnil, 1950.—Afrotropical: D.R. Congo, Kenya, Lesotho, Malawi, Rwanda, South Africa, Tanzania, Uganda, Zimbabwe.
Carcelia peraequalis Mesnil, 1950b: 18. Syntypes, males and females (possibly 1 male in CNC [O’Hara 1996: 152], 1 male in IRSNB). Type locality: Zimbabwe, Harare [as “Salisbury”].
persimilis Mesnil, 1950.—Afrotropical: Madagascar, South Africa.
Carcelia persimilis Mesnil, 1950b: 17. Lectotype male (MNHN), by fixation of O’Hara (1996: 153) (treatment of a male labelled as “TYPE” from Fort-Dauphin in MNHN as the holotype is regarded as a lectotype fixation). Type locality: Madagascar, Toliara, Tôlanaro [also commonly known as Taolagnaro or Fort Dauphin and published as “Fort-Dauphin”].
Note: O’Hara (1996: 153) accepted the specimen labelled as “TYPE” in MNHN as the holotype of Carcelia persimilis Mesnil, 1950 under the assumption the species was likely described from a single specimen. However, Mesnil’s description (1950b: 17) clearly mentions both sexes, thus indicating syntypes. O’Hara’s (1996: 153) treatment of the “TYPE” in MNHN as the holotype of C. persimilis is regarded as a lectotype fixation.
vaga (Curran, 1927).—Afrotropical: Uganda.
Zenillia vaga Curran, 1927d: 332. Holotype male (BMNH). Type locality: Uganda, Kampala.
vara (Curran, 1927).—Afrotropical: Ghana, Kenya, South Africa, Tanzania.
Zenillia vara Curran, 1927d: 331. Holotype male (BMNH). Type locality: Kenya, Kabete [ca. 1°16′S 36°43′E, near Nairobi].
vexor (Curran, 1927).—Afrotropical: South Africa.
Zenillia vexor Curran, 1927d: 330. Holotype male (SANC). Type locality: South Africa, KwaZulu-Natal, Durban.
Subgenus EURYCLEA Robineau-Desvoidy, 1863
EURYCLEA Robineau-Desvoidy, 1863a: 290. Type species: Euryclea tibialis Robineau-Desvoidy, 1863, by original designation [Palaearctic].
setifrons Mesnil, 1949.—Afrotropical: D.R. Congo, Nigeria (new record, CNC), Uganda.
Carcelia (Eucarcelia) setifrons Mesnil, 1949a: 90. Holotype male (MRAC). Type locality: D.R. Congo, Katanga, Lubumbashi [as “Elisabethville”].
Possibly undescribed spp.: Yemen, as “Carcelia (Caricelia) sp. 1 cf. vexor”, “Carcelia (Caricelia) sp. 2”, and “Carcelia (Caricelia) sp. 3” (Zeegers 2007: 378).
Genus CARCELIATHRIX Cerretti & O’Hara, gen. n.
CARCELIATHRIX Cerretti & O’Hara, gen. n. Type species: Phorocera crassipalpis Villeneuve, 1938, by present designation.
Note: This new genus is described in the New Taxa of Afrotropical Tachinidae section.
crassipalpis (Villeneuve, 1938).—Afrotropical: D.R. Congo. Comb. n.
Phorocera crassipalpis Villeneuve, 1938c: 2. Lectotype male (MRAC), by designation herein (see Lectotype Designations section). Type locality: D.R. Congo, Équateur, Bomputu.
claripalpis. Incorrect subsequent spelling of crassipalpis Villeneuve, 1938 (original usage not found but spelling listed by Crosskey 1980b: 867).
Note: Crosskey (1980b: 867) treated Phorocera crassipalpis Villeneuve, 1938 as an unplaced species in the “Carceliini”.
Undescribed sp. 1: Namibia (NNIC, examined by PC).
Undescribed sp. 2: South Africa (NMB, examined by PC).
Genus CESTONIA Rondani, 1861
CESTONIA Rondani, 1861b: 105. Type species: Cestonia cineraria Rondani, 1861, by monotypy [Palaearctic].
canariensis Villeneuve, 1936.—Afrotropical: U.A. Emirates. Palaearctic: N. Africa (Canary Is.), M. East (Israel).
Cestonia canariensis Villeneuve in Frey, 1936: 145. Syntypes, 1 male and 1 female (FMNHH). Type locality: Canary Islands, Gran Canaria, Las Palmas de Gran Canaria.
Note: Cestonia canariensis Villeneuve, 1936, was redescribed by Herting (1981: 3) from the original syntypes.
harteni Zeegers, 2007.—Afrotropical: Yemen.
Cestonia harteni Zeegers, 2007: 381. Holotype female (RMNH). Type locality: Yemen, Suq Bani Mansour (15°05′15″N 43°52′10″E).
Note: Zeegers (2010: 677) recognized “Cestonia cf. harteni Zeegers” from U.A. Emirates.
rufipes Zeegers, 2007.—Afrotropical: Yemen.
Cestonia rufipes Zeegers, 2007: 382. Holotype male (RMNH). Type locality: Yemen, Al Kawd [as “Al Kowd”] (15°14′52″N 43°15′16″E).
rutilans Villeneuve, 1929.—Afrotropical: Senegal, Yemen. Palaearctic: N. Africa (NE. Africa).
Cestonia rutilans Villeneuve, 1929a: 102. Syntypes, 1 male and 1 female (not located). Type locality: Egypt, Al Qāhirah [as “Caire”].
Genus CESTONIONERVA Villeneuve, 1929
CESTONIONERVA Villeneuve, 1929b: 43. Type species: Conogaster petiolata Villeneuve, 1910, by subsequent designation of Townsend (1936b: 137).
Note: Cestonionerva Villeneuve, 1929 was “Formé pour Conogaster petiolata Villen.” (Villeneuve 1929b: 43) and a new species of the genus was also described in the same paper. Crosskey (1980b: 876) and Herting and Dely-Draskovits (1993: 223) interpreted Conogaster petiolata as the type species of Cestonionerva by original designation. However, a fixation by original designation requires an explicit designation of a type species (Article 68.2 of the Code, ICZN 1999), which is lacking in this instance. The type species of Cestonionerva Villeneuve, 1929 was therefore fixed later by the subsequent designation of Townsend (1936b: 137).
petiolata (Villeneuve, 1910).—Afrotropical: U.A. Emirates, Yemen. Palaearctic: C. Asia, M. East (Israel), Mongolia, N. Africa (Canary Is., NE. Africa), Pal. China.
Conogaster petiolata Villeneuve in Becker, 1910b: 144 [also 1910b: 14]. Holotype female (NHMW). Type locality: Yemen, Suquţrá [as “Sokótra”].
Genus CHRYSERYCIA Mesnil, 1977
CHRYSERYCIA Mesnil, 1977b: 185. Type species: Chryserycia fulviceps Mesnil, 1977, by original designation.
fulviceps Mesnil, 1977.—Afrotropical: Madagascar.
Chryserycia fulviceps Mesnil, 1977b: 186. Holotype female (MNHN). Type locality: Madagascar, Antsiranana, Montagne d’Ambre [Parc National, ca. 12°36′S 49°8′E].
Genus DESCAMPSINA Mesnil, 1956
DESCAMPSINA Mesnil, 1956b: 76. Type species: Descampsina sesamiae Mesnil, 1956, by original designation.
sesamiae Mesnil, 1956.—Afrotropical: Cameroon, D.R. Congo (new record, IRSNB [PC]), Nigeria.
Descampsina sesamiae Mesnil, 1956b: 76. Holotype, unspecified sex [male, examined by PC] (MNHN). Type locality: Cameroon, Garoua.
Note: Mesnil (1956b: 76–77) described Descampsina sesamiae from both sexes from Garoua (Cameroon) and wrote “Type dans ma collection”, without giving the sex. O’Hara (1996: 156) treated the type series as comprising syntypes in CNC and MNHN but this is incorrect because a holotype (as “Type”) was designated in the original description.
Genus DIAPROCHAETA Mesnil, 1970
DIAPROCHAETA Mesnil, 1970b: 103. Type species: Diaprochaeta (Diaprochaeta) illustris Mesnil, 1970, by original designation.
illustris Mesnil, 1970.—Afrotropical: Zimbabwe.
Diaprochaeta (Diaprochaeta) illustris Mesnil, 1970b: 105. Holotype male (CNC). Type locality: Zimbabwe, “Sankishya” [not located].
Genus DRINO Robineau-Desvoidy, 1863
Subgenus DRINO Robineau-Desvoidy, 1863
DRINO Robineau-Desvoidy, 1863a: 250. Type species: Drino volucris Robineau-Desvoidy, 1863 (= Tachina lota Meigen, 1824), by original designation [Palaearctic].
STURMIODORIA Townsend, 1928: 391. Type species: Sturmiodoria facialis Townsend, 1928, by original designation.
cordata (Curran, 1927).—Afrotropical: Burundi, D.R. Congo, Guinea, Malawi, Rwanda.
Sturmia cordata Curran, 1927a: 12. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
Note: Sturmia cordata Curran, 1927 is moved here from Drino subgenus Palexorista Townsend, 1921 based on examination of the holotype by PC.
facialis (Townsend, 1928).—Afrotropical: D.R. Congo. Palaearctic: Pal. China. Oriental: India, Indonesia, Malaysia, Orien. China, Philippines, Sri Lanka, Taiwan, Thailand.
Sturmiodoria facialis Townsend, 1928: 392. Holotype female (USNM). Type locality: Philippines, Basilan.
Note: Sturmiodoria facialis Townsend, 1928 was recorded from Africa (D.R. Congo) by Verbeke (1962b: 51) but Crosskey (1984: 284) commented that “confirmation of identity in Africa [is] needed”.
lota (Meigen, 1824).—Afrotropical: Tanzania. Palaearctic: Europe (all except SW. Eur., Turkey), Japan, Pal. China [Ningxia], Russia (W. Russia, W. Siberia, S. Far East). Oriental: Orien. China.
Tachina lota Meigen, 1824: 326. Lectotype male (MNHN), by designation of Herting (1972: 9). Type locality: not given (Europe).
Note: Tachina lota Meigen, 1824 was recorded from Africa (Tanzania) by Mesnil (1959: 8) but Crosskey (1984: 284) commented that “confirmation of identity in Africa [is] needed”.
Subgenus PALEXORISTA Townsend, 1921
PALEXORISTA Townsend, 1921: 134. Type species: Tachina succini Giebel, 1862 (as “Tichina succini Giebel”), by monotypy.
PROSTURMIA Townsend, 1927c: 69. Type species: Prosturmia profana Townsend, 1927 (= Masicera solennis Walker, 1858), by original designation [Oriental].
PROSTURMINA Mesnil, 1949b: 103 (as subgenus of Drino Robineau-Desvoidy, 1863). Nomen nudum (proposed after 1930 without designation of type species; no included species).
PROSTURMINA Mesnil, 1949c: 8, 32 (as subgenus of Drino Robineau-Desvoidy, 1863). Nomen nudum (proposed after 1930 without type designation from three included species).
PROSTURMINA Mesnil, 1951: 161 (as subgenus of Drino Robineau-Desvoidy, 1863). Nomen nudum (proposed after 1930 without type designation; no included species).
PROSTURMINA Mesnil, 1970b: 110 (as subgenus of Drino Robineau-Desvoidy, 1863). Type species: Sturmia vigilans Villeneuve, 1933 (= Sturmia pulchra Curran, 1927), by original designation.
Note: The nomenclatural history of Prosturmina Mesnil was discussed by O’Hara (1996: 128) and Evenhuis and O’Hara (2008: 67).
amicula Mesnil, 1949.—Afrotropical: Cameroon, Ghana, Mozambique, Nigeria, Senegal, Tanzania.
Drino (Prosturmia) amicula Mesnil, 1949c: 30. Syntypes, males and females (1 male in CNC, 2 males in MNHN). Type localities: Mozambique (Rio Zambeze [Tambara according to label data, Cooper and O’Hara 1996: 30; ca. 16°43′S 34°15′E]) and Senegal (Bambey).
ampliceps (Karsch, 1886).—Afrotropical: Angola.
Masicera (Blepharipa) ampliceps Karsch, 1886b: 340. Holotype, unspecified sex [female, examined by JEOH] (ZMHB). Type locality: Angola, Pungo Andongo.
aureocincta Mesnil, 1977.—Afrotropical: Madagascar.
Drino (Prosturmia) aureocincta Mesnil, 1977b: 179. Holotype male (MNHN). Type locality: Madagascar, Toliara, Sakaraha.
aureola Mesnil, 1970.—Afrotropical: Sierra Leone.
Drino (Prosturmina) aureola Mesnil, 1970b: 110. Holotype male (CNC). Type locality: Sierra Leone, Bafodia [as “Bafodea”, ca. 9°41′N 11°43′E].
aurifera (Villeneuve, 1943).—Afrotropical: D.R. Congo.
Sturmia aurifera Villeneuve, 1943a: 36. Syntypes, males and females (2 males in CNC). Type localities: D.R. Congo, Équateur, Eala and Maniema, Lubutu.
crassiseta Mesnil, 1968.—Afrotropical: South Africa.
Drino crassiseta Mesnil, 1968b: 5. Holotype male (SMNS). Type locality: South Africa, Western Cape, Cape Town, Kirstenbosch.
curvipalpis (van der Wulp, 1893).—Misidentification, not Afrotropical [known from Palaearctic, Oriental and Australasian regions].
Note: An unknown species was recorded as “Drino (Prosturmia T.T.) unisetosa Bar.” (originally described as Sturmia (Sturmia) unisetosa Baranov, 1932, currently a synonym of Drino curvipalpis (van der Wulp, 1893)) from Tanzania by Mesnil (1959: 7). Misidentification (Crosskey 1980b: 872).
flavicans (Wiedemann, 1819).—Afrotropical: D.R. Congo, Malawi, South Africa, Uganda.
Tachina flavicans Wiedemann, 1819: 24. Type(s), female (not located). Type locality: South Africa, Western Cape, Cape of Good Hope [as “Prom. bon. sp.” = “Promontorium Bonae Spei”].
Sturmia congolensis Villeneuve, 1910a: 253. Syntypes, 3 females (not located). Type locality: D.R. Congo (as “Congo”, p. 249).
Note: Villeneuve (1910a) described four species from “Congo”. Curran (1927f: 122) treated one of them (Sturmia aureiventris Villeneuve, 1910) as described from D.R. Congo (as “Belgian Congo”), and used “Belgian Congo” and “Congo” interchangeably in this work and some others. We think it likely that Villeneuve (1910a), like Curran, used “Congo” in the sense of present-day D.R. Congo. However, Crosskey (1980b) interpreted Villeneuve’s Congo as the present-day country of Congo.
flaviseta (Thomson, 1869).—Afrotropical: Mauritius.
Masicera flaviseta Thomson, 1869: 522. Type(s), unspecified sex (NHRS). Type locality: Mauritius.
gilva (Hartig, 1838).—Misidentification, not Afrotropical [known from Palaearctic Region].
Note: An unknown species was recorded as “Sturmia gilva Hartig” (originally described as Tachina gilva Hartig, 1838) from D.R. Congo by Curran (1927f: 116, 1928b: 393). Misidentifications (not recorded from the Afrotropical Region by Herting and Dely-Draskovits 1993: 207).
gilvoides (Curran, 1927).—Afrotropical: D.R. Congo, South Africa.
Sturmia gilvoides Curran, 1927f: 117. Holotype male (SANC). Type locality: South Africa, Mpumalanga, Barberton.
idonea (Brauer & Bergenstamm, 1891).—Afrotropical: ?Eritrea, Mozambique, South Africa.
Argyrophylax idonea Brauer & Bergenstamm, 1891: 344 [also 1891: 40]. Type(s), male (NHMW, not located by JEOH). Type locality: South Africa, Western Cape, Cape of Good Hope [as “Cap b. sp.” = “Cap Bonae Spei”].
Sturmia partitor Curran, 1927f: 116. Holotype male (SANC). Type locality: South Africa, Free State, Bloemfontein.
Note: Bezzi’s (1908b: 30) record of Drino idonea (Brauer & Bergenstamm, 1891) (as Sturmia (Argyrophylax) idonea) from Eritrea needs confirmation.
imberbis (Wiedemann, 1830).—Afrotropical: D.R. Congo, Kenya, Malawi, South Africa, Sudan, Tanzania, U.A. Emirates, Uganda, Yemen. Palaearctic: C. Asia, Europe (SC. Eur., Turkey), M. East (all), N. Africa (Canary Is., NE. Africa), Transcaucasia.
Tachina imberbis Wiedemann, 1830: 317. Syntypes, 2 or more males (lost, Crosskey 1967b: 93, Ziegler 2011: 8). Type locality: Egypt.
Sturmia zonata Curran, 1927d: 336. Holotype male (BMNH). Type locality: Uganda, Entebbe.
Note: See Herting (1984: 193, note 142) and Ziegler (2011: 7–8) for a discussion of the identities of Tachina imberbis Wiedemann, 1830, Sturmia zonata Curran, 1927, and Phorcida latigena Mesnil, 1944. Zeegers (2007: 386, 2010: 679) treated T. imberbis as a nomen dubium and used S. zonata as the valid name for this taxon.
inconspicua (Meigen, 1830).—Misidentification, not Afrotropical [known from Palaearctic and Oriental regions].
Note: An unknown species was recorded as “Sturmia (Sturmia) inconspicua” (originally described as Tachina inconspicua Meigen, 1830) from Tanzania by Speiser (1910: 140). The same or a similar species was recorded from Malawi and Uganda by Villeneuve (1913c: 29, as “Sturmia inconspicua”), from D.R. Congo and South Africa by Curran (1927f: 118, 1928b: 393, as “Sturmia inconspicua”) and from Seychelles by Barraclough (2009: 304, as “Drino inconspicua” but noting “Confirmation of identity required”). It was also recorded as “Sturmia bimaculata” (originally described as Tachina bimaculata Hartig, 1838, currently a synonym of Drino inconspicua (Meigen)) from D.R. Congo by Curran (1927f: 118, 1928b: 394). Misidentifications (Crosskey 1980b: 871, 872).
iterata Mesnil, 1949.—Afrotropical: South Africa, Uganda.
Drino (Prosturmia) iterata Mesnil, 1949c: 31. Syntypes, males and females (1 male in CNC). Type localities: Uganda and South Africa.
latigena (Mesnil, 1944).—Afrotropical: Djibouti, U.A. Emirates. Palaearctic: M. East (Israel), N. Africa (NE. Africa).
Phorcida latigena Mesnil, 1944: 15. Holotype male (MNHN). Type locality: Djibouti, Obock [as “Obok”].
Tachina imberbis of authors (e.g., Crosskey 1967b: 93–94, 1980b: 872 [in part], as “Palexorista imberbis”), not Wiedemann, 1830. Misidentification (Herting 1984: 193, note 142; Ziegler 2011: 7–8).
lavinia (Curran, 1927).—Afrotropical: D.R. Congo, Uganda.
Sturmia lavinia Curran, 1927c: 14. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
laxa (Curran, 1927).—Afrotropical: Botswana, Kenya, Malawi, South Africa, Sudan, Swaziland, Tanzania, Uganda, Zimbabwe. Oriental: India.
Sturmia laxa Curran, 1927d: 335. Holotype male (BMNH). Type locality: Tanzania, Morogoro.
mayneana (Villeneuve, 1930).—Afrotropical: D.R. Congo.
Sturmia mayneana Villeneuve, 1930b: 59. Syntypes, males and females (“plusieurs individus”) (MRAC). Type locality: D.R. Congo, Équateur, Eala.
melancholica Mesnil, 1949.—Afrotropical: Zimbabwe.
Drino (Prosturmia) melancholica Mesnil, 1949c: 16. Syntypes, 1 male and 1 female (CNC). Type locality: Zimbabwe, Harare [as “Salisbury”].
nova Mesnil, 1949.—Afrotropical: Madagascar.
Drino (Prosturmia) nova Mesnil, 1949c: 27. Syntypes, males and females (1 male in CNC, MNHN). Type locality: Madagascar, Toliara, Bekily.
obliterata Mesnil, 1949.—Afrotropical: Malawi, Senegal, South Africa.
Drino (Prosturmia) patruelis obliterata Mesnil, 1949c: 18. Syntypes, males and females (MNHN). Type localities: Malawi (Mt. Mulanje [as “Mt. Mlanje”]), Senegal (Bambey), and South Africa.
parachrysops (Bezzi, 1925).—Afrotropical: Ghana, Kenya, Mali, Nigeria, Senegal, Yemen. Palaearctic: M. East (M. East [Saudi Arabia, Dawah 2011: 5]). Oriental: India, ?Indonesia, Malaysia, Sri Lanka.
Sturmia parachrysops Bezzi, 1925b: 114. Lectotype male (BMNH), by designation of Crosskey (1967b: 78). Type locality: Malaysia, Peninsular Malaysia, Kuala Lumpar.
patruelis Mesnil, 1949.—Afrotropical: Malawi, South Africa, Tanzania, Uganda, Zimbabwe.
Drino (Prosturmia) patruelis Mesnil, 1949c: 17. Syntypes, males and probably females (“nombreux exemplaires”) (1 male and possibly other syntypes in CNC). Type localities: South Africa and Zimbabwe (Harare [as “Salisbury”]).
pulchra (Curran, 1927).—Afrotropical: D.R. Congo, Uganda.
Sturmia pulchra Curran, 1927a: 16. Holotype male (BMNH). Type locality: Uganda, Entebbe.
Sturmia vigilans Villeneuve, 1933: 278. Holotype female (MRAC). Type locality: D.R. Congo, Équateur, Eala.
quadrizonula (Thomson, 1869).—Afrotropical: widespread, including D.R. Congo, Ghana, Kenya, Saint Helena, São Tomé & Principe, Senegal, Seychelles, South Africa, Tanzania, Uganda, Zimbabwe (Crosskey 1977: 152, in part).
Masicera quadrizonula Thomson, 1869: 521. Lectotype female (NHRS), by designation of Crosskey (1970: 580). Type locality: Saint Helena.
Note: Masicera quadrizonula Thomson, 1869 was redescribed by Crosskey (1970: 580, 1977: 151).
rufa Zeegers, 2007.—Afrotropical: Yemen.
Drino rufa Zeegers, 2007: 385. Holotype male (RMNH). Type locality: Yemen, Sana’a (15°21′17″N 44°12′24″E).
salva (Wiedemann, 1830).—Afrotropical: South Africa.
Tachina salva Wiedemann, 1830: 340. Type(s), female (1 syntype in ZMUC, Zimsen 1954: 23). Type locality: South Africa [as “China”, in error according to Crosskey 1980b: 872].
subaurata (Walker, 1853).—Afrotropical: Madagascar, South Africa.
Tachina subaurata Walker, 1853: 298. Type(s) female (BMNH). Type locality: South Africa, Western Cape, Cape of Good Hope [as “Cape”].
succini (Giebel, 1862).—Afrotropical: ?Tanzania.
Tachina succini Giebel, 1862: 319. Holotype female (NMCL). Type locality: not given (in copal; “East Africa presumed”, Crosskey 1980b: 872).
Note: Tachina succini Giebel, 1862 was described from a copal inclusion originally thought to be an amber fossil (Crosskey 1966c: 133). Its provenance is unknown but likely East Africa, particularly Zanzibar, a popular source of copal since the early 1800s.
tenella (Bezzi, 1911).—Afrotropical: South Africa.
Erycia (Bactromyia) tenella Bezzi, 1911: 60. Holotype female (USNM). Type locality: South Africa, Gauteng, Pretoria.
terrosa Mesnil, 1949.—Afrotropical: Madagascar.
Drino (Prosturmia) terrosa Mesnil, 1949c: 20. Type(s), unspecified sex (MNHN). Type locality: Madagascar, Toliara, Bekily.
ugandana (Curran, 1927).—Afrotropical: Burundi, D.R. Congo, Malawi, South Africa, Uganda, Zimbabwe.
Sturmia ugandana Curran, 1927c: 16. Holotype male (AMNH; not BMNH, see Arnaud and Owen 1981: 239). Type locality: Uganda, Entebbe.
Subgenus ZYGOBOTHRIA Mik, 1891
ZYGOBOTHRIA Mik, 1891: 193. Type species: Sturmia atropivora Robineau-Desvoidy, 1830, by original designation.
FORMOSODORIA Townsend, 1933: 475. Type species: Sturmia (Argyrophylax) dilabida Villeneuve, 1916 (= Meigenia ciliata van der Wulp, 1881), by original designation.
atropivora (Robineau-Desvoidy, 1830).—Afrotropical: “widespread Afrotrop Reg.” (Crosskey 1980b: 874), including D.R. Congo, Ghana, Kenya, Madagascar, Malawi, Mauritius, Mozambique, Namibia, Sierra Leone, South Africa, Tanzania, U.A. Emirates, Uganda. Palaearctic: C. Asia, Europe (all except British Is., Scand.), Japan, M. East (all), N. Africa (Canary Is., NW. Africa), Pal. China, Russia (W. Russia), Transcaucasia. Oriental: India, Indonesia, Laos, Malaysia, Orien. China, Ryukyu Is., Sri Lanka. Australasian: Australia.
Sturmia atropivora Robineau-Desvoidy, 1830: 171. Syntypes, more than 80 males and females (lost, Herting 1974a: 24). Type locality: not given (France).
Sturmia masakensis Curran, 1927f: 117. Holotype male (BMNH). Type locality: Uganda, Masaka.
masakesnsis. Incorrect subsequent spelling of masakensis Curran, 1927 (Curran 1928b: 388).
ciliata (van der Wulp, 1881).—Afrotropical: “widespread mainland Afrotrop. Reg.” (Crosskey 1980b: 874), including Ghana, Malawi, South Africa, U.A. Emirates, Uganda. Palaearctic: Pal. China. Oriental: India, Indonesia, Sri Lanka, Taiwan. Australasian: Australia, N. Australasian.
Meigenia ciliata van der Wulp, 1881: 38. Lectotype male (RMNH), by designation of Crosskey (1967c: 104). Type locality: Indonesia, Sumatera, Alahanpanjang [as “Alahan pandjang”].
Sturmia (Argyrophylax) dilabida Villeneuve, 1916c: 479. Type(s), unspecified number and including at least 1 male (SAMC, not located by JEOH). Type locality: South Africa, KwaZulu-Natal, Durban.
Note: Sturmia dilabida Villeneuve, 1916 was described from one or more specimens, at least one of which was male. The type locality was given as Durban and the depository as SAMC. There are several specimens in SAMC identified by Villeneuve as S. dilabida but none from Durban. Unless type material of S. dilabida is discovered in SAMC or is proven to have existed there, Townsend’s (1941: 270) mention of “Ht male” from Durban in SAMC cannot be accepted as a lectotype fixation. The identity of S. dilabida was confused with that of Sturmia convergens (Wiedemann, 1824) by Villeneuve (1933: 280). Townsend (1932: 32, 1933: 475) erred in citing the type locality of S. dilabida as Taiwan [as “Formosa”] and the type depository as SDEI [as “Berlin-Dahlem”]. Later, Townsend (1941: 270) correctly cited the type locality as Durban.
Sturmia munroi Curran, 1927c: 17. Holotype male (SANC). Type locality: South Africa, Eastern Cape, East London.
Sturmia (Sturmia) macrophallus Baranov, 1932: 76. Lectotype male (SDEI), by designation of Crosskey (1967c: 105). Type locality: Taiwan, P’ingtung Hsien, Changkou [as “Kankau”, near Hengch’un].
Formosodoria foeda Villeneuve, 1933: 280 (as “Formosodoria foeda T. T.”). Nomen nudum (proposed in synonymy [with Sturmia dilabida Villeneuve, 1916 and Tachina convergens Wiedemann, 1824, the latter misidentified] and not made available by subsequent usage before 1961).
Tachina convergens of Mesnil (1951: 169, as “Drino convergens”), not Wiedemann, 1824. Misidentification (Crosskey 1963: 77, 1980b: 874).
grandicornis Mesnil, 1977.—Afrotropical: Madagascar.
Drino (Zygobothria) grandicornis Mesnil, 1977b: 178. Holotype male (MNHN). Type locality: Madagascar, Fianarantsoa, Mananjary.
Genus EUGAEDIOXENIS Cerretti, O’Hara & Stireman, 2015
EUGAEDIOXENIS Cerretti, O’Hara & Stireman inCerretti et al., 2015: 494. Type species: Gaedioxenis haematodes Villeneuve, 1937, by original designation.
haematodes (Villeneuve, 1937).—Afrotropical: South Africa.
Gaedioxenis haematodes Villeneuve, 1937a: 207. Holotype male (CNC). Type locality: South Africa, “Colonie du Cap” ([former Cape Province], between Somerset West and Strand according to label data, Cooper and O’Hara 1996: 39).
horridus Cerretti, O’Hara & Stireman, 2015.—Afrotropical: South Africa.
Eugaedioxenis horridus Cerretti, O’Hara & Stireman inCerretti et al., 2015: 501. Holotype male (MZUR). Type locality: South Africa, Western Cape, Anysberg Nature Reserve, 840m (33°26′37.76″S 20°47′29.25″E).
Genus HYPERSARA Villeneuve, 1935
HYPERSARA Villeneuve, 1935a: 139. Type species: Hypersara argentata Villeneuve, 1935, by monotypy.
argentata Villeneuve, 1935.—Afrotropical: D.R. Congo, Nigeria.
Hypersara argentata Villeneuve, 1935a: 140. Holotype male (not located). Type locality: D.R. Congo, Nord-Kivu, Walikale [ca. 1°25′S 28°00′E].
Undescribed sp.: Ethiopia (TAU, examined by PC).
Genus INTRAPALES Villeneuve, 1938
INTRAPALES Villeneuve, 1938c: 8. Type species: Intrapales remotella Villeneuve, 1938, by monotypy.
hirsuta Mesnil, 1977.—Afrotropical: Madagascar.
Intrapales hirsuta Mesnil, 1977b: 185. Holotype male (MNHN). Type locality: Madagascar, Toamasina, Périnet [ca. 18°55′S 48°25′E].
insularis Mesnil, 1977.—Afrotropical: Madagascar.
Intrapales insularis Mesnil, 1977b: 184. Holotype male (MNHN). Type locality: Madagascar, Fianarantsoa, Anosimparihy [ca. 21°30′S 47°59′E].
remotella Villeneuve, 1938.—Afrotropical: D.R. Congo, Nigeria (new record, CNC), Sierra Leone, Tanzania.
Intrapales remotella Villeneuve, 1938c: 8. Syntypes, 2 males and 1 female (IRSNB). Type locality: D.R. Congo, Équateur, Eala.
Genus KAISERIOLA Mesnil, 1970
KAISERIOLA Mesnil, 1970b: 105 (as subgenus of Diaprochaeta Mesnil, 1970). Type species: Diaprochaeta (Kaiseriola) aperta Mesnil, 1970, by original designation.
Note: Kaiseriola Mesnil, 1970 was treated as a synonym of Diaprochaeta Mesnil, 1970 by Crosskey (1980b: 877) but was later recognized as a genus by Crosskey (1984: 201, 294).
aperta (Mesnil, 1970).—Afrotropical: Mozambique (new record, JOS [PC]), South Africa.
Diaprochaeta (Kaiseriola) aperta Mesnil, 1970b: 105. Holotype male (CNC). Type locality: South Africa, KwaZulu-Natal, Durban.
obscura (Mesnil, 1970).—Afrotropical: Madagascar.
Diaprochaeta (Kaiseriola) obscura Mesnil, 1970b: 106. Holotype male (MNHN). Type locality: Madagascar, Toamasina, Moramanga.
Genus LUBUTANA Villeneuve, 1938
LUBUTANA Villeneuve, 1938c: 10. Type species: Lubutana divaricata Villeneuve, 1938, by original designation.
divaricata Villeneuve, 1938.—Afrotropical: D.R. Congo, Ethiopia, Ghana, Malawi, Nigeria, Sierra Leone, Uganda.
Lubutana divaricata Villeneuve, 1938c: 10. Syntypes, males (IRSNB). Type localities: D.R. Congo (Maniema, Lubutu; Nord-Kivu, Walikale [ca. 1°25′S 28°00′E]), Malawi (Mt. Mulanje [as “Mont Mlanje”]) and Nigeria (Degema).
mayeri Mesnil, 1955.—Afrotropical: Nigeria.
Lubutana mayeri Mesnil, 1955: 363. Holotype female (CNC). Type locality: Nigeria, Oshogbo.
perplexa Mesnil, 1955.—Afrotropical: D.R. Congo, Rwanda, Uganda.
Lubutana perplexa Mesnil, 1955: 362. Holotype female (MRAC). Type locality: Rwanda, eastern foothills of Volcan Muhabura [as “Muhavura”], 2100m [ca. 1°23′S 29°44′E].
Genus LYDELLA Robineau-Desvoidy, 1830
LYDELLA Robineau-Desvoidy, 1830: 112. Type species: Lydella grisescens Robineau-Desvoidy, 1830, by subsequent designation of Robineau-Desvoidy (1863a: 855) [Palaearctic].
METOPOSISYROPS Townsend, 1916d: 320. Type species: Metoposisyrops oryzae Townsend, 1916, by original designation [Oriental].
Note: Metoposisyrops Townsend, 1916 was synonymized with Lydella Robineau-Desvoidy, 1830 by Woodley (1994: 135).
sesamiae (Mesnil, 1968).—Afrotropical: D.R. Congo (new record, IRSNB [PC]), Mozambique (new record, MZUR [PC]), Namibia (new record, MZUR [PC]), Nigeria, Uganda.
Metagonistylum sesamiae Mesnil, 1968b: 4. Holotype female (CNC). Type locality: Uganda, Kidetok, Serere [ca. 1°30′N 33°33′E].
Genus MADREMYIA Townsend, 1916
MADREMYIA Townsend, 1916d: 305. Type species: Madremyia parva Townsend, 1916 (= Phorocera saundersii Williston, 1889), by original designation [Neotropical]. New record.
Note: Madremyia Townsend, 1916 is newly recorded from the Afrotropical Region for a species previously placed in Phryxe Robineau-Desvoidy, 1830.
setinervis (Mesnil, 1968).—Afrotropical: Tanzania. Comb. n.
Phryxe setinervis Mesnil, 1968b: 5. Holotype female (SMNS). Type locality: Tanzania, southwest side of Mt. Kilimanjaro [as “Kilimandjaro”], 3500m.
Note: Phryxe setinervis Mesnil, 1968 was treated as a species of Phryxe Robineau-Desvoidy, 1830 by Crosskey (1980b: 879) but is moved here to Madremyia Townsend, 1916.
Genus MYXARCHICLOPS Villeneuve, 1916
MYXARCHICLOPS Villeneuve, 1916c: 494. Type species: Myxarchiclops caffer Villeneuve, 1916, by subsequent designation of Townsend (1936b: 222).
caffer Villeneuve, 1916.—Afrotropical: South Africa.
Myxarchiclops caffer Villeneuve, 1916c: 495. Lectotype male (CNC), by designation herein (see Lectotype Designations section). Type locality: South Africa, Western Cape, Cape Town.
major Villeneuve, 1930.—Afrotropical: South Africa.
Myxarchiclops major Villeneuve, 1930a: 353. Syntypes, 2 females (CNC). Type locality: South Africa, “Colonie du Cap” ([former Cape Province], Somerset West according to label data, Cooper and O’Hara 1996: 54).
Genus NEOLYDELLA Mesnil, 1939
NEOLYDELLA Mesnil, 1939a: 209 (as subgenus of Lydella Robineau-Desvoidy, 1830). Type species: Lydella (Neolydella) pruinosa Mesnil, 1939, by monotypy.
pruinosa (Mesnil, 1939).—Afrotropical: Madagascar.
Lydella (Neolydella) pruinosa Mesnil, 1939a: 209. Syntypes, 3 males (MNHN). Type locality: Madagascar, Toliara, Bekily, “région sud de l’Ile”.
Genus NILEA Robineau-Desvoidy, 1863
NILEA Robineau-Desvoidy, 1863a: 275. Type species: Nilea innoxia Robineau-Desvoidy, 1863, by original designation [Palaearctic].
longicauda (Mesnil, 1970).—Afrotropical: Madagascar. Comb. n.
Sturmia longicauda Mesnil, 1970b: 91. Holotype male (MNHN). Type locality: Madagascar, Toamasina, Moramanga.
Note: Sturmia longicauda Mesnil, 1970 was treated as a species of Sturmia Robineau-Desvoidy, 1830 by Crosskey (1980b: 874) but is moved here to Nilea Robineau-Desvoidy, 1863.
perplexa Mesnil, 1977.—Afrotropical: Burundi (new record, MZUR [PC]), Madagascar, Mozambique (new record, MZUR [PC]), South Africa (new record, NMDA [PC]).
Nilea perplexa Mesnil, 1977b: 188. Holotype male (MNHN). Type locality: Madagascar, Toamasina, Foulpointe [ca. 17°41′S 49°31′E].
Undescribed sp.: Tanzania (TAU, examined by PC).
Genus PARADRINO Mesnil, 1949
PARADRINO Mesnil, 1949b: 103 (as subgenus of Drino Robineau-Desvoidy, 1863). Type species: Sturmia halli Curran, 1939 (as “Paradrino Halli Curr.”, p. 100), by monotypy (see Evenhuis and O’Hara 2008: 66).
halli (Curran, 1939).—Afrotropical: Botswana, Tanzania, Uganda, Zimbabwe.
Sturmia halli Curran, 1939: 2. Holotype male (AMNH). Type locality: Zimbabwe, Kadoma [as “Gatooma”].
Sturmia rhodesiensis Jones, 1939: 16. Syntypes, males and females (BMNH). Type locality: Zimbabwe, Mazoe.
Note: Jones (1939: 15) wrote in a footnote on the first page of his paper: “After the present manuscript had been sent to the printers, Curran published a description of this species under the name of Sturmia halli sp. n. (1939, Amer. Mus. Nov. 1022, pp. 2–3).” Since the name Sturmia rhodesiensis was not explicitly proposed in synonymy with Sturmia halli, it is treated as both an available name and a subjective synonym of S. halli.
Undescribed species of “?Paradrino”: Yemen (Zeegers 2007: 392).
Genus PHRYXE Robineau-Desvoidy, 1830
PHRYXE Robineau-Desvoidy, 1830: 158. Type species: Phryxe athaliae Robineau-Desvoidy, 1830 (= Tachina vulgaris Fallén, 1810), by subsequent designation of Robineau-Desvoidy (1863a: 329, 358) (as vulgaris, with athaliae in synonymy) [Palaearctic].
Note: The single species recognized in Phryxe Robineau-Desvoidy, 1830 by Crosskey (1980b: 879), Phryxe setinervis Mesnil, 1968, is moved herein to Madremyia Townsend, 1916. We record Phryxe in the Afrotropical Region from an undescribed species.
Undescribed sp.: Ethiopia (TAU, examined by PC).
Genus PSEUDOPERICHAETA Brauer & Bergenstamm, 1889
PSEUDOPERICHAETA Brauer & Bergenstamm, 1889: 92 [also 1890: 24]. Type species: Pseudoperichaeta major Brauer & Bergenstamm, 1889 (= Phryxe palesioidea Robineau-Desvoidy, 1830), by monotypy [Palaearctic].
ACHAETONEURILLA Mesnil, 1939a: 210 (as subgenus of Pseudoperichaeta Brauer & Bergenstamm, 1889). Type species: Pseudoperichaeta (Achaetoneurilla) madecassa Mesnil, 1939, by monotypy.
laevis Villeneuve, 1932.—Afrotropical: Nigeria, Tanzania, Uganda, Zimbabwe.
Pseudoperichaeta laevis Villeneuve, 1932: 285. Syntypes, males and females (not located). Type locality: Zimbabwe, Harare [as “Salisbury”].
Phorocera bolyodes Curran, 1933: 166. Holotype female (BMNH). Type locality: Zimbabwe, Harare [as “Salisbury”].
bothyodes. Incorrect subsequent spelling of bolyodes Curran, 1933 (original usage not found but spelling listed by Crosskey 1980b: 880).
leo (Curran, 1941).—Afrotropical: Zimbabwe.
Phorocera leo Curran, 1941: 10. Holotype female (AMNH). Type locality: Zimbabwe, Mutare [as “Umtali”].
Pseudoperichaeta pilosa Villeneuve, 1942a: 52. Syntypes, 2 males (1 male in CNC). Type locality: Zimbabwe, Hurungwe [as “Urungwe”], Gota Gota.
madecassa Mesnil, 1939.—Afrotropical: Madagascar.
Pseudoperichaeta (Achaetoneurilla) madecassa Mesnil, 1939a: 210. Syntypes, 12 males and females (3 males and 2 females in CNC, MNHN). Type locality: Madagascar, Toliara, Bekily, “région sud de l’Ile”.
nestor (Curran, 1927).—Afrotropical: D.R. Congo, Nigeria, Tanzania.
Phorocera nestor Curran, 1927c: 12. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
pacta Villeneuve, 1932.—Afrotropical: D.R. Congo, Mauritius, South Africa, Zimbabwe.
Pseudoperichaeta pacta Villeneuve, 1932: 285. Holotype female (not located). Type locality: South Africa, Western Cape, “région de Cape-Town”.
sallax (Curran, 1927).—Afrotropical: D.R. Congo.
Phorocera sallax Curran, 1927c: 11. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
Genus PTILOCATAGONIA Mesnil, 1956
PTILOCATAGONIA Mesnil, 1956b: 79 (as subgenus of Sisyropa Brauer & Bergenstamm, 1889). Type species: Sisyropa (Ptilocatagonia) viridescens Mesnil, 1956, by monotypy.
viridescens (Mesnil, 1956).—Afrotropical: Sierra Leone, Tanzania, Zambia.
Sisyropa (Ptilocatagonia) viridescens Mesnil, 1956b: 79. Holotype male (SMNS). Type locality: Tanzania, Msingi [ca. 4°20′S 34°34′E].
Note: Mesnil (1956b: 79) described Sisyropa viridescens from a “Mâle capturé à Msingi (Ruwenzori)”. The type locality of Msingi is in Tanzania, whereas “Ruwenzori” refers to the Rwenzori Range on the border between D.R. Congo and Uganda. The country of the type locality was incorrectly cited as Uganda by Crosskey (1980b: 873) and O’Hara (1996: 160).
Genus SENOMETOPIA Macquart, 1834
SENOMETOPIA Macquart, 1834: 160 [also 1834: 296]. Type species: Carcelia aurifrons Robineau-Desvoidy, 1830 (= Tachina excisa Fallén, 1820), by subsequent designation of Townsend (1916b: 8) (earlier type fixations set aside by ICZN 2012: 242; see Evenhuis and Thompson 1990: 237 and O’Hara and Evenhuis 2011: 61) [Palaearctic].
STENOMETOPIA Agassiz, 1846b: 351. Unjustified emendation of Senometopia Macquart, 1834.
EOCARCELIA Townsend, 1919b: 582. Type species: Eocarcelia ceylanica Townsend, 1919, by original designation [Oriental].
EOCARCELIOPSIS Townsend, 1928: 392. Type species: Eocarceliopsis bakeri Townsend, 1928, by original designation [Oriental].
EUCARCELIA Baranov, 1934: 393. Type species: Tachina excisa Fallén, 1820, by original designation [Palaearctic].
albatella (Villeneuve, 1941).—Afrotropical: D.R. Congo, Malawi.
Carcelia albatella Villeneuve, 1941b: 125. Syntypes, 1 male and 1 female (MRAC). Type locality: D.R. Congo, Sud-Kivu, Kalembelembe to Baraka.
evolans (Wiedemann, 1830).—Afrotropical: Côte d’Ivoire, Senegal, Sierra Leone, ?Yemen.
Tachina evolans Wiedemann, 1830: 321. Type(s), unspecified sex (not located). Type locality: Sierra Leone.
Note: Tachina evolans Wiedemann, 1830 has been misidentified from other places in the Afrotropical Region and from the Palaearctic Region, as noted by Crosskey (1980b: 865), Shima (2006: 64, 66) and O’Hara et al. (2009: 78). Given such a history of misidentifications, we treat the record from Yemen by Zeegers (2007: 396) as questionable. Curran (1927d: 327) examined the “type” of T. evolans but did not state where he had seen it or give any details about it.
hectica (Speiser, 1910).—Afrotropical: Kenya, Tanzania, Uganda.
Carcelia hectica Speiser, 1910: 141. Holotype male (NHRS). Type locality: Tanzania, Mt. Kilimanjaro [as “Kilimandjaro”], valley at Kibongoto [as “Kibonoto”].
illota (Curran, 1927).—Afrotropical: Nigeria, South Africa, Tanzania. Oriental: India, Laos, Orien. China. Australasian: Australia.
Zenillia illota Curran, 1927d: 328. Holotype male (BMNH). Type locality: Tanzania, Morogoro.
judicabilis (Mesnil, 1949).—Afrotropical: D.R. Congo, Malawi, Zimbabwe.
Carcelia (Eucarcelia) evolans judicabilis Mesnil, 1949a: 90. Holotype, unspecified sex [male, examined by PC] (MRAC). Type locality: D.R. Congo, Katanga, Lubumbashi [as “Elisabethville”].
laetifica (Mesnil, 1949).—Afrotropical: D.R. Congo, Ghana, Nigeria.
Carcelia (Eucarcelia) evolans laetifica Mesnil, 1949a: 89. Holotype male (MRAC). Type locality: D.R. Congo, Katanga, Lubumbashi [as “Elisabethville”].
norma (Curran, 1927).—Afrotropical: Malawi, Tanzania, Uganda.
Zenillia norma Curran, 1927d: 329. Holotype male (BMNH). Type locality: Uganda, Bugoma Forest [ca. 1°16′N 30°57′E].
Genus SISYROPA Brauer & Bergenstamm, 1889
SISYROPA Brauer & Bergenstamm, 1889: 163 [also 1890: 95]. Type species: Tachina thermophila Wiedemann, 1830, by monotypy [Oriental].
STYLURODORIA Townsend, 1933: 476. Type species: Stylurodoria stylata Townsend, 1933, by original designation.
CTENOPHOROCEROPSIS Baranov, 1938: 408. Type species: Ctenophoroceropsis yerburyi Baranov, 1938, by original designation.
POUJADEA Mesnil, 1949b: 102. Nomen nudum (proposed after 1930 without designation of type species; no included species) (see Evenhuis and O’Hara 2008: 67).
EOCATAGONIA Mesnil, 1949b: 103 (as subgenus Sisyropa Brauer & Bergenstamm, 1889). Nomen nudum (proposed after 1930 without designation of type species; no included species) (see Evenhuis and O’Hara 2008: 66).
POUJADEA Mesnil, 1950c: 108. Type species: Zenillia insolita Curran, 1927, by monotypy (see Evenhuis and O’Hara 2008: 67).
EOCATAGONIA Mesnil, 1950c: 148 (as subgenus Sisyropa Brauer & Bergenstamm, 1889). Type species: Sisyropa (Eocatagonia) argyrata Mesnil, 1950, by monotypy (see Evenhuis and O’Hara 2008: 66).
argyrata Mesnil, 1950.—Afrotropical: Senegal.
Sisyropa (Eocatagonia) argyrata Mesnil, 1950c: 148. Holotype male (MNHN). Type locality: Senegal.
boveyi Mesnil, 1958.—Afrotropical: Ghana, Guinea, Kenya, Nigeria, Tanzania.
Sisyropa (Catagonia) boveyi Mesnil, 1958: 252. Holotype male (ETHZ). Type locality: Guinea, Réserve de la Biosphère des Monts Nimba [as “Réserve du Mt Nimba”], foot of Mont Nimba.
insolita (Curran, 1927).—Afrotropical: D.R. Congo.
Zenillia insolita Curran, 1927c: 5. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
insoleta. Incorrect subsequent spelling of insolita Curran, 1927 (Curran 1927d: 327).
madecassa Mesnil, 1944.—Afrotropical: Madagascar.
Sisyropa formosa madecassa Mesnil, 1944: 14. Holotype male (MNHN). Type locality: Madagascar, Fianarantsoa, Ikongo-Ankarimbelo region, Forêt Tanala.
negator (Curran, 1927).—Afrotropical: D.R. Congo.
Sturmia negator Curran, 1927c: 15. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
negastor. Incorrect subsequent spelling of negator Curran, 1927 (Curran 1928b: 391).
stylata (Townsend, 1933).—Afrotropical: Ghana, Mali, Nigeria, Sierra Leone, Sudan. Oriental: India, Sri Lanka, Taiwan.
Stylurodoria stylata Townsend, 1933: 476. Holotype female (SDEI). Type locality: Taiwan, P’ingtung Hsien, Changkou [as “Kankau”, near Hengch’un].
subdistincta (Villeneuve, 1916).—Afrotropical: Côte d’Ivoire, Ethiopia, Ghana, Senegal, South Africa, Tanzania.
Catagonia subdistincta Villeneuve, 1916c: 484. Syntypes, 2 males (SAMC, not located by JEOH). Type locality: South Africa, KwaZulu-Natal, Durban.
Sisyropa cinerosa Mesnil, 1944: 15. Holotype male (MNHN). Type locality: Senegal, Bambey.
yerburyi (Baranov, 1938).—Afrotropical: Yemen.
Ctenophoroceropsis yerburyi Baranov, 1938: 409. Holotype male (BMNH, Sabrosky and Crosskey 1969: 39). Type locality: Yemen, ‘Adan [as “Aden”].
yerburi. Incorrect subsequent spelling of yerburyi Baranov, 1938 (original usage not found but spelling listed by Crosskey 1980b: 873).
Possibly undescribed spp.: Nigeria (BMNH, Crosskey 1984: 281).
Genus STURMIOPSIS Townsend, 1916
STURMIOPSIS Townsend, 1916d: 313. Type species: Sturmiopsis inferens Townsend, 1916, by original designation.
RHODESINA Curran, 1939: 3 (junior homonym of Rhodesina Malloch, 1921). Type species: Rhodesina parasitica Curran, 1939, by original designation.
CURRANOMYIA Townsend in Cuthbertson & Munro, 1941: 115 (nomen novum for Rhodesina Curran, 1939).
Note: Barraclough (2004) published a review of Sturmiopsis Townsend, 1916 but was apparently unaware of the key to species of Sturmiopsis and the description of Sturmiopsis setifrons Mesnil, 1977 by Mesnil (1977b: 186–187).
inferens Townsend, 1916.—Afrotropical: Madagascar (probably introduced, Barraclough 2004: 12). Oriental: Bangladesh, Bhutan, India, Indonesia, Malaysia, Orien. China, Nepal, Philippines.
Sturmiopsis inferens Townsend, 1916d: 313. Holotype female (USNM). Type locality: Indonesia, Jawa, Bogor [as “Buitenzorg”].
parasitica (Curran, 1939).—Afrotropical: Benin, Ghana, Kenya, Nigeria, Senegal, Tanzania, Zimbabwe. Oriental: India (introduced, Barraclough 2004: 17).
Rhodesina parasitica Curran, 1939: 3. Holotype male (AMNH). Type locality: Zimbabwe, Harare [as “Salisbury”].
Sturmiopsis angustifrons Mesnil, 1959: 11. Holotype male (SMNS). Type locality: Tanzania, Kisangara.
setifrons Mesnil, 1977.—Afrotropical: Madagascar.
Sturmiopsis setifrons Mesnil, 1977b: 187. Holotype male (MNHN). Type locality: Madagascar, Fianarantsoa, Ambalavao.
Genus STYLOCARCELIA Zeegers, 2007
STYLOCARCELIA Zeegers, 2007: 396. Type species: Stylocarcelia stylata Zeegers, 2007, by original designation.
stylata Zeegers, 2007.—Afrotropical: Yemen.
Stylocarcelia stylata Zeegers, 2007: 396. Holotype male (RMNH). Type locality: Yemen, Sana’a (15°21′17″N 44°12′24″E).
Genus THECOCARCELIA Townsend, 1933
THECOCARCELIA Townsend, 1933: 471. Type species: Argyrophylax pelmatoprocta Brauer & Bergenstamm, 1891 (= Masicera acutangulata Macquart, 1851), by original designation [Palaearctic].
THELYCARCELIA Townsend, 1933: 475. Type species: Thelycarcelia thrix Townsend, 1933 (= Sturmia sumatrana Baranov, 1932), by original designation [Oriental].
Note: The relative priority of Thecocarcelia Townsend, 1933 and Thelycarcelia Townsend, 1933, when the two are treated as synonyms, was established by Mesnil (1950b: 20), as the First Reviser (Article 24.2.2 of the Code, ICZN 1999).
acutangulata (Macquart, 1851).—Afrotropical: “W. Afr. to E. Afr. & sthn Afr.” (Crosskey 1980b: 866), including D.R. Congo, Madagascar. Palaearctic: Europe (all except Scand., Turkey), Japan, Transcaucasia.
Masicera acutangulata Macquart, 1851a: 478. Type(s), female (MHNL or lost). Type locality: Switzerland, Chur [as “Coire”].
Masicera incedens Rondani, 1861b: 22. Type(s), female (MZF, Herting 1969: 195; 1 female syntype and 1 male non-type in MZF [examined by PC]). Type locality: Italy, plain near Parma.
Argyrophylax pelmatoprocta Brauer & Bergenstamm, 1891: 344 [also 1891: 40]. Syntypes, males and females (2 males and 4 females in NHMW). Type locality: “M.-Europa”.
Note: Argyrophylax pelmatoprocta Brauer & Bergenstamm, 1891 was described from an unspecified number of males and females from “M.-Europa”. Herting (1974b: 140) reported on a female syntype from Bisamberg [near Wien, Austria] in NHMW but this collection includes an additional two males and three females identified as pelmatoprocta by “B. B.” or “Bergenst.” from other localities in Europe and these specimens are considered syntypes as well (examined by JEOH).
ebenina Mesnil, 1950.—Afrotropical: D.R. Congo, South Africa.
Thecocarcelia ebenina Mesnil, 1950b: 21. Syntypes, males and possibly females (not located). Type locality: South Africa, KwaZulu-Natal.
flavicosta Zeegers, 2007.—Afrotropical: Yemen.
Thecocarcelia flavicosta Zeegers, 2007: 398. Holotype male (RMNH). Type locality: Yemen, Laḩij [as “Lahj”] (13°03′28″N 44°53′02″E).
latifrons Mesnil, 1949.—Afrotropical: Mozambique, South Africa, Uganda, Zimbabwe.
Thecocarcelia latifrons Mesnil, 1949b: 56. Holotype male (CNC). Type locality: Mozambique, Rio Zambezi, near Chemba, “Nova Choupanga”.
Note: Zeegers (2010: 681) recognized “Thecocarcelia cf. latifrons Mesnil” from U.A. Emirates.
latimana Mesnil, 1950.—Afrotropical: South Africa.
Thecocarcelia latimana Mesnil, 1950b: 22. Syntypes, males and females (not located). Type locality: South Africa.
pauciseta Mesnil, 1977.—Afrotropical: Madagascar.
Thecocarcelia pauciseta Mesnil, 1977b: 181. Holotype male (NHMB [“to be returned to MNHN”, O’Hara 1996: 152]). Type locality: Madagascar, Toamasina, 15km [south of] Mananara, Ivontaka [ca. 16°18′S 49°49′E].
robusta Mesnil, 1950.—Afrotropical: D.R. Congo.
Thecocarcelia robusta Mesnil, 1950b: 22. Syntypes, males (1 male in CNC). Type locality: D.R. Congo, Équateur, Eala.
trichops Herting, 1967.—Afrotropical: South Africa, Zambia. Palaearctic: Europe (W. Eur., SW. Eur., SC. Eur., SE. Eur.), Japan, Pal. China.
Thecocarcelia trichops Herting, 1967: 4. Holotype male (CNC). Type locality: France, Vaucluse, Lagnes.
Note: Specimens from the Afrotropical Region identified as Thecocarcelia trichops Herting, 1967 should be checked to confirm their identity.
ventralis Mesnil, 1959.—Afrotropical: D.R. Congo, Ghana, Nigeria, Sierra Leone, Tanzania.
Thecocarcelia ventralis Mesnil, 1959: 2. Holotype male (SMNS). Type locality: Tanzania, “Torina” [not located].
vibrissata Mesnil, 1977.—Afrotropical: Madagascar.
Thecocarcelia vibrissata Mesnil, 1977b: 181. Holotype male (MNHN). Type locality: Madagascar, Fianarantsoa, Ifanadiana [ca. 21°18′S 47°38′E].
Genus THELAIRODRINO Mesnil, 1954
THELAIRODRINO Mesnil, 1954c: 470 (as subgenus of Thelairosoma Villeneuve, 1916). Type species: Thelairosoma gracilis Mesnil, 1952, by original designation [Oriental].
anaphe (Curran, 1927).—Afrotropical: Cameroon, D.R. Congo, Kenya, Malawi, Nigeria, Tanzania, Zimbabwe.
Sturmia anaphe Curran, 1927e: 447. Holotype male (BMNH). Type locality: Tanzania, Morogoro.
Note: Sturmia anaphe, described by Curran (1927e: 447), was referred to as “Sturmia anaphe, sp. n.” by Curran (1927f: 126) with the accompanying note, “It will be described fully in the Entomologische Mitteilungen” (i.e., in Curran 1927e, which was published first).
cardinalis (Mesnil, 1949).—Afrotropical: D.R. Congo.
Drino cardinalis Mesnil, 1949a: 91. Holotype, unspecified sex [male, examined by PC] (MRAC). Type locality: D.R. Congo, Katanga, Lubumbashi [as “Elisabethville”].
potina (Curran, 1927).—Afrotropical: South Africa.
Sturmia potina Curran, 1927f: 118. Holotype male (SANC). Type locality: South Africa, KwaZulu-Natal, Port Shepstone.
Genus THELAIROSOMA Villeneuve, 1916
THELAIROSOMA Villeneuve, 1916c: 499. Type species: Thelairosoma fumosum Villeneuve, 1916, by monotypy.
SEYRIGOMYIA Mesnil, 1944: 11. Type species: Seyrigomyia fulvella Mesnil, 1944, by original designation.
LESPESIOPSIS Mesnil, 1954c: 471 (as subgenus of Thelairosoma Villeneuve, 1916). Type species: Thelairosoma (Lespesiopsis) coerulescens Mesnil, 1954, by monotypy.
THELAIROXENIS Mesnil, 1954c: 472 (as subgenus of Thelairosoma Villeneuve, 1916). Type species: Thelairosoma (Thelairoxenis) pallidum Mesnil, 1954, by original designation.
angustifrons (Villeneuve, 1916).—Afrotropical: D.R. Congo, Malawi, Mozambique, Nigeria, South Africa, Tanzania, Uganda.
Sturmia (Blepharipoda) angustifrons Villeneuve, 1916c: 478. Syntypes, 3 males and 1 female (1 male in SAMC). Type locality: South Africa, KwaZulu-Natal, Durban.
atrum Mesnil, 1970.—Afrotropical: Madagascar.
Thelairosoma (Thelairosoma) atrum Mesnil, 1970b: 101. Holotype male (MNHN). Type locality: Madagascar, Toamasina, Périnet [ca. 18°55′S 48°25′E].
brunnescens (Villeneuve, 1934).—Afrotropical: Rwanda, Uganda.
Erycia brunnescens Villeneuve, 1934d: 69. Lectotype female (IRSNB), by designation herein (see Lectotype Designations section). Type locality: Uganda, Rwenzori Range [as “Ruwenzori”], 2500m.
carbonatum (Mesnil, 1944).—Afrotropical: Madagascar.
Seyrigomyia carbonata Mesnil, 1944: 13. Holotype male (MNHN). Type locality: Madagascar, Toliara, Bekily.
coerulescens Mesnil, 1954.—Afrotropical: Burundi, D.R. Congo, Rwanda, Tanzania.
Thelairosoma (Lespesiopsis) coerulescens Mesnil, 1954c: 471. Holotype male (CNC). Type locality: northwest Tanzania, edge of virgin forest, 1800–2200m.
comatum Villeneuve, 1938.—Afrotropical: Uganda.
Thelairosoma comatum Villeneuve, 1938b: 2. Holotype male (IRSNB). Type locality: Uganda, Rwenzori Range [as “Ruwenzori”], 2300m.
diaphanum Mesnil, 1954.—Afrotropical: D.R. Congo.
Thelairosoma (Thelairoxenis) diaphanum Mesnil, 1954c: 472. Holotype male (IRSNB). Type locality: D.R. Congo, Équateur, Eala.
flavipalpe Villeneuve, 1938.—Afrotropical: D.R. Congo.
Thelairosoma flavipalpe Villeneuve, 1938b: 3. Holotype male (IRSNB). Type locality: D.R. Congo, Nord-Kivu, Walikale [ca. 1°25′S 28°00′E].
fulvellum (Mesnil, 1944).—Afrotropical: Madagascar.
Seyrigomyia fulvella Mesnil, 1944: 12. Holotype, unspecified sex (MNHN). Type locality: Madagascar, Toliara, Bekily.
fumosum Villeneuve, 1916c: 500.—Afrotropical: D.R. Congo, Ghana, Malawi, Mozambique (new record, MZUR [PC]), South Africa, Tanzania.
Thelairosoma fumosum Villeneuve, 1916c: 500. Lectotype male (SAMC), by fixation of Townsend (1940: 98) (mention of “Ht male” from Durban in SAMC is regarded as a lectotype fixation for the male syntype in SAMC labelled by Villeneuve as “Typ.” [examined by JEOH]). Type locality: South Africa, KwaZulu-Natal, Durban.
hybridum Mesnil, 1970.—Afrotropical: Madagascar.
Thelairosoma (Seyrigomyia) hybrida Mesnil, 1970b: 103. Holotype male (MNHN). Type locality: Madagascar, Antananarivo, Antananarivo [as “Tananarive”].
ingrami Mesnil, 1970.—Afrotropical: Uganda.
Thelairosoma (Seyrigomyia) ingrami Mesnil, 1970b: 103. Holotype male (CNC). Type locality: Uganda, Serere [ca. 1°30′N 33°33′E].
longicorne Mesnil, 1954.—Afrotropical: Zimbabwe.
Thelairosoma (Thelairoxenis) longicorne Mesnil, 1954c: 473. Holotype male (BMNH). Type locality: Zimbabwe, Harare [as “Salisbury”].
lutescens Mesnil, 1954.—Afrotropical: Malawi, South Africa, Zimbabwe.
Thelairosoma (Seyrigomyia) lutescens Mesnil, 1954c: 474. Holotype, unspecified sex (BMNH, not located by D. Whitmore, pers. comm.). Type locality: South Africa.
major Mesnil, 1970.—Afrotropical: Madagascar.
Thelairosoma (Seyrigomyia) major Mesnil, 1970b: 102. Holotype male (MNHN). Type locality: Madagascar, Antananarivo, Mandraka [near Antananarivo, not located].
melancholicum Mesnil, 1970.—Afrotropical: Madagascar.
Thelairosoma (Seyrigomyia) melancholica Mesnil, 1970b: 102. Holotype male (MNHN). Type locality: Madagascar, Toamasina, Périnet [ca. 18°55′S 48°25′E].
obversum Villeneuve, 1943.—Afrotropical: Zimbabwe.
Thelairosoma obversum Villeneuve, 1943a: 40. Syntypes, 3 males (not located). Type locality: Zimbabwe, Harare [as “Salisbury”].
pallidum Mesnil, 1954.—Afrotropical: D.R. Congo, Malawi, Nigeria.
Thelairosoma (Thelairoxenis) pallidum Mesnil, 1954c: 472. Holotype male (MRAC). Type locality: D.R. Congo, Katanga, Lubumbashi [as “Elisabethville”].
palposum Villeneuve, 1938.—Afrotropical: “W. Afr. to E. Afr. & sthn Afr.” (Crosskey 1980b: 881), including D.R. Congo, Gabon.
Thelairosoma palposum Villeneuve, 1938b: 2. Syntypes, 1 male and 1 female (1 male in IRSNB). Type localities: D.R. Congo, Nord-Kivu, Walikale [ca. 1°25′S 28°00′E] and Gabon, “Bas-Ogooué” [delta region of the Rivière Ogooué].
pulchellum (Mesnil, 1944).—Afrotropical: Madagascar.
Seyrigomyia pulchella Mesnil, 1944: 13. Holotype, unspecified sex (MNHN). Type locality: Madagascar, central plateau of Fianarantsoa.
quadriguttatum (Mesnil, 1944).—Afrotropical: Madagascar.
Seyrigomyia quadriguttata Mesnil, 1944: 12. Holotype, unspecified sex [male, see O’Hara 1996: 154] (MNHN). Type locality: Madagascar.
rosatum Villeneuve, 1943.—Afrotropical: Malawi.
Thelairosoma rosatum Villeneuve, 1943a: 39. Holotype female (not located). Type locality: Malawi, Mt. Mulanje [as “Mt. Mlanje”].
triste Mesnil, 1970.—Afrotropical: Madagascar.
Thelairosoma (Seyrigomyia) tristis Mesnil, 1970b: 102. Holotype male (CNC). Type locality: Madagascar, Toamasina, Périnet [ca. 18°55′S 48°25′E].
varipes Villeneuve, 1943.—Afrotropical: Malawi.
Thelairosoma varipes Villeneuve, 1943a: 39. Syntypes, 3 males and 4 females (not located). Type locality: Malawi.
Genus THELYCONYCHIA Brauer & Bergenstamm, 1889
THELYCONYCHIA Brauer & Bergenstamm, 1889: 89 [also 1890: 21]. Type species: Masicera (Ceromasia) solivaga Rondani, 1861, by monotypy.
TORINAMYIA Mesnil, 1959: 2. Type species: Torinamyia delicatula Mesnil, 1959, by monotypy.
delicatula (Mesnil, 1959).—Afrotropical: Tanzania, Uganda.
Torinamyia delicatula Mesnil, 1959: 2. Holotype male (SMNS). Type locality: Tanzania, “Torina” [not located].
solivaga (Rondani, 1861).—Afrotropical: Botswana, U.A. Emirates, Yemen. Palaearctic: C. Asia, Europe (all except British Is., Scand.), Japan, M. East (Israel), Pal. China, Russia (E. Siberia, S. Far East), Transcaucasia. Oriental: Pakistan.
Masicera (Ceromasia) solivaga Rondani, 1861b: 24. Type(s), male (MZF, Herting 1969: 201; 8 male syntypes and 6 female non-types in MZF [examined by PC]). Type locality: Italy, plain near Parma.
Note: Thelyconychia solivaga (Rondani, 1861) of current authors is likely a species complex but is treated here as a single species pending further study. Crosskey’s (1980b: 867) record of T. solivaga from Canary Islands may have been based on a misidentification because the species was not recorded from there by Tschorsnig and Báez (2002) or Tschorsnig et al. (2004).
Genus THELYMYIOPS Mesnil, 1950
THELYMYIOPS Mesnil, 1950b: 10 (as subgenus of Carcelia Robineau-Desvoidy, 1830). Type species: Carcelia coniformis Villeneuve, 1941, by monotypy. Status n.
Note: Thelymyiops Mesnil, 1950 was treated as a subgenus of Carcelia Robineau-Desvoidy, 1830 by Crosskey (1980b: 866). It is here raised to a genus and the characters that distinguish it will be given in the Tachinidae chapter of the Manual of Afrotropical Diptera (in prep.).
coniformis (Villeneuve, 1941).—Afrotropical: D.R. Congo, Ghana, Tanzania, Uganda.
Carcelia coniformis Villeneuve, 1941b: 124. Holotype female (IRSNB). Type locality: D.R. Congo, Équateur, Eala.
Unplaced species of Eryciini
varicornis Curran, 1940.—Afrotropical: Zambia, Zimbabwe.
Phorocera varicornis Curran, 1940: 7. Holotype female (AMNH). Type locality: border between Zambia and Zimbabwe, Victoria Falls.
Note: Phorocera varicornis Curran, 1940 was treated as an “Unplaced species of Goniinae” [= Exoristinae] by Crosskey (1980b: 881) and is moved here based on examination of the holotype by PC. It cannot be placed to genus at the present time.
Tribe ETHILLINI
Genus AMNONIA Kugler, 1971
AMNONIA Kugler, 1971: 71. Type species: Amnonia carmelitana Kugler, 1971, by original designation.
carmelitana Kugler, 1971.—Afrotropical: Ethiopia (new record, TAU [PC]), Kenya (new record, TAU [PC]). Palaearctic: M. East (Israel).
Amnonia carmelitana Kugler, 1971: 71. Holotype male (TAU). Type locality: Israel, Zikhron Ya’aqov.
Note: Amnonia carmelitana Kugler, 1971 is newly recorded from the Afrotropical Region.
deemingi Zeegers, 2010.—Afrotropical: U.A. Emirates.
Amnonia deemingi Zeegers, 2010: 674. Holotype male (RMNH). Type locality: U.A. Emirates, 7km south of Jazīrat al Hamrā [as “al-Jazirat al-Hamra”] (25°39′N 55°45′E).
Genus CALLIETHILLA Shima, 1979
CALLIETHILLA Shima, 1979: 147. Type species: Calliethilla caerulea Shima, 1979, by original designation [Oriental].
hirta Cerretti, 2012.—Afrotropical: Uganda.
Calliethilla hirta Cerretti, 2012: 322. Holotype male (TAU). Type locality: Uganda, Rwenzori Range [as “Ruwenzori”], Itojo.
Genus ETHILLA Robineau-Desvoidy, 1863
ETHILLA Robineau-Desvoidy, 1863a: 202. Type species: Tachina aemula Meigen, 1824, by original designation [Palaearctic].
ETHYLLA Mesnil, 1939d: 32. Unjustified emendation of Ethilla Robineau-Desvoidy, 1863 (see Evenhuis et al. 2010: 76).
adiscalis Mesnil, 1977.—Afrotropical: Madagascar.
Ethilla adiscalis Mesnil, 1977b: 173. Holotype male (MNHN). Type locality: Madagascar, Antananarivo, Manjakatompo [ca. 19°21′S 47°18′E].
tenor (Curran, 1927).—Afrotropical: ?Angola, D.R. Congo, ?Kenya, ?Malawi.
Zenillia tenor Curran, 1927c: 5. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
Note: Crosskey (1984: 270) recorded specimens in BMNH from Angola, Kenya and Malawi that are “probably” Zenillia tenor Curran, 1927.
Possibly undescribed sp.: South Africa (BMNH, Crosskey 1984: 270).
Genus ETHYLLOIDES Verbeke, 1970
ETHYLLOIDES Verbeke, 1970: 286. Type species: Ethylloides emdeni Verbeke, 1970, by original designation.
emdeni Verbeke, 1970.—Afrotropical: South Africa.
Ethylloides emdeni Verbeke, 1970: 288. Holotype male (MZLU). Type locality: South Africa, Western Cape, Cape Peninsula, Hout Bay, Skoorsteenkop.
Genus GYNANDROMYIA Bezzi, 1923
GYNANDROMYIA Bezzi, 1923: 97. Type species: Gynandromyia seychellensis Bezzi, 1923, by original designation.
ZENILLIANA Curran, 1927c: 3 (as subgenus of Zenillia Robineau-Desvoidy, 1830). Type species: Zenillia (Zenilliana) devastator Curran, 1927 (= Myxexorista habilis Brauer & Bergenstamm, 1891), by monotypy.
ZELINDOMYIA Verbeke, 1962a: 166. Type species: Zelindomyia grossa Verbeke, 1962, by original designation.
TRYPHEROSOMA Verbeke, 1962a: 167. Type species: Trypherosoma gilva Verbeke, 1962, by original designation.
Note: Trypherosoma Verbeke, 1962 and Zelindomyia Verbeke, 1962 were synonymized with Gynandromyia Bezzi, 1923 by Crosskey (1984: 200, 271).
bafwankei Verbeke, 1962.—Afrotropical: D.R. Congo.
Gynandromyia bafwankei Verbeke, 1962a: 172. Lectotype male (IRSNB), by fixation of Verbeke (1962b: 44) (examination of “Type, 1♂” from Bafwankei is regarded as a lectotype fixation). Type locality: D.R. Congo, Orientale, Bafwakei [as “Bafwankei”, ca. 1°41′N 27°02′E, near Bomili].
Note: The name Gynandromyia bafwankei was made available by Verbeke (1962a: 172) in a key that was apparently intended to precede the full description by Verbeke (1962b: 43, as “Gynandromyia bafwankei n. sp.”). No specimens were mentioned in the first work but two males, “Type” and “Paratype”, were cited in the second. We regard the “Type” as the lectotype of G. bafwankei by fixation of Verbeke (1962b: 44).
basilewskyi (Verbeke, 1960).—Afrotropical: Tanzania.
Zenilliana basilewskyi Verbeke, 1960: 337. Holotype male (MRAC). Type locality: Tanzania, Olkokola, Mt. Meru, towards northwest, 2500–2600m.
crypta (Verbeke, 1962).—Afrotropical: D.R. Congo.
Trypherosoma crypta Verbeke, 1962a: 167, 168. Holotype male (IRSNB). Type locality: D.R. Congo, Orientale, Bafwakei [as “Bafwankei”, ca. 1°41′N 27°02′E, near Bomili].
fumigata (Verbeke, 1962).—Afrotropical: D.R. Congo.
Trypherosoma fumigata Verbeke, 1962a: 167, 168. Holotype male (IRSNB). Type locality: D.R. Congo, Équateur, Eala.
gilva (Verbeke, 1962).—Afrotropical: D.R. Congo.
Trypherosoma gilva Verbeke, 1962a: 167, 168. Holotype male (IRSNB). Type locality: D.R. Congo, Équateur, Eala.
grossa (Verbeke, 1962).—Afrotropical: D.R. Congo.
Zelindomyia grossa Verbeke, 1962a: 167. Holotype male (IRSNB). Type locality: D.R. Congo, Orientale, Mapolo.
habilis (Brauer & Bergenstamm, 1891).—Afrotropical: “widespread W. Afr., E. Afr. & sthn Afr.” (Crosskey 1980b: 861), including D.R. Congo, Malawi, South Africa.
Myxexorista habilis Brauer & Bergenstamm, 1891: 332 [also 1891: 28] (as “habilis Wd. litt. Coll. Wiedm.”). Type(s), male (NHMW, not located by JEOH). Type locality: South Africa, Western Cape, Cape of Good Hope [as “Cap b. sp.” = “Cap Bonae Spei”].
Zenillia (Zenilliana) devastator Curran, 1927c: 3. Holotype female (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
Zenillia fuscicosta Curran, 1927c: 4. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
invaginata (Villeneuve, 1939).—Afrotropical: D.R. Congo.
Zenilliana devastator invaginata Villeneuve, 1939: 9. Syntypes, 2 females (not located). Type localities: D.R. Congo, Orientale, Bafwakei [as “Bafwankei”, ca. 1°41′N 27°02′E, near Bomili] and Équateur, Irebu.
kibatiana Verbeke, 1962.—Afrotropical: D.R. Congo.
Gynandromyia kibatiana Verbeke, 1962a: 172. Lectotype male (IRSNB), by fixation of Verbeke (1962b: 41) (examination of “Type, 1♂” from Kibati is regarded as a lectotype fixation). Type locality: D.R. Congo, Nord-Kivu, Parc National des Virunga [as “P.N.A”, former Parc National Albert], foot of Mt. Nyiragongo [as “Nyaragongo”], Kibati [ca. 1°36′S 29°16′E].
Note: The name Gynandromyia kibatiana was made available by Verbeke (1962a: 172) in a key that was apparently intended to precede the full description by Verbeke (1962b: 39, as “Gynandromyia kibatiana n. sp.”). No specimens were mentioned in the first work but a male “Type” and two “Paratypes” (one male and one female) were cited in the second. We regard the “Type” as the lectotype of G. kibatiana by fixation of Verbeke (1962b: 41).
mesnili Verbeke, 1962.—Afrotropical: Burundi.
Gynandromyia mesnili Verbeke, 1962a: 172. Holotype male (MRAC). Type locality: Burundi, Bururi, 1800–2000m.
Note: The name Gynandromyia mesnili was made available by Verbeke (1962a: 172) in a key that was apparently intended to precede the full description by Verbeke (1962b: 38, as “Gynandromyia mesnili n. sp.”). No specimens were mentioned in the first work but a single specimen, a male “Type”, was cited in the second. Since the nominal species was clearly based on a single specimen, we regard the “Type” as the holotype of G. mesnili by monotypy in Verbeke (1962a).
prima Verbeke, 1962.—Afrotropical: Ghana, Kenya, Malawi, South Africa, Uganda, Zimbabwe.
Gynandromyia prima Verbeke, 1962a: 172. Lectotype male (IRSNB), by fixation of Verbeke (1962b: 37) (examination of “Type, 1♂” from Aburi is regarded as a lectotype fixation). Type locality: Ghana, Aburi.
Note: The name Gynandromyia prima was made available by Verbeke (1962a: 172) in a key that was apparently intended to precede the full description by Verbeke (1962b: 36, as “Gynandromyia prima n. sp.”). No specimens were mentioned in the first work but a male “Type” and a series of “Paratypes” (of both sexes) were cited in the second. We regard the “Type” as the lectotype of G. prima by fixation of Verbeke (1962b: 37).
saegeri Verbeke, 1962.—Afrotropical: D.R. Congo.
Gynandromyia saegeri Verbeke, 1962a: 171. Holotype male (MRAC). Type locality: D.R. Congo, Orientale, Parc National de la Garamba [as “P.N.G.”].
Note: The name Gynandromyia saegeri was made available by Verbeke (1962a: 171) in a key that was apparently intended to precede the full description by Verbeke (1962b: 44, as “Gynandromyia saegeri n. sp.”). No specimens were mentioned in the first work but a single specimen, a male “Type”, was cited in the second. Since the nominal species was clearly based on a single specimen, we regard the “Type” as the holotype of G. saegeri by monotypy in Verbeke (1962a).
seychellensis Bezzi, 1923.—Afrotropical: Seychelles.
Gynandromyia seychellensis Bezzi, 1923: 98. Holotype female [not male as published, Crosskey 1984: 272] (BMNH). Type locality: Seychelles, Mahé Is., Cascade Estate, ca. 1000ft.
Genus MYCTEROMYIELLA Mesnil, 1966
MYCTEROMYIA Mesnil, 1949b: 102. Nomen nudum (proposed after 1930 without designation of type species; no included species) (see Evenhuis and O’Hara 2008: 66).
MYCTEROMYIA Mesnil, 1950c: 107 (junior homonym of Mycteromyia Philippi, 1865). Type species: Mycteromyia laetifica Mesnil, 1950, by monotypy (see Evenhuis and O’Hara 2008: 66) [Australasian].
MYCTEROMYIELLA Mesnil, 1966: 232 (nomen novum for Mycteromyia Mesnil, 1950).
Undescribed sp.: Angola (BMNH, Crosskey 1980b: 862, Crosskey 1984: 269).
Genus NEMORILLOIDES Brauer & Bergenstamm, 1891
NEMORILLOIDES Brauer & Bergenstamm, 1891: 355 [also 1891: 51]. Type species: Nemorilloides flaviventris Brauer & Bergenstamm, 1891, by monotypy.
carbonata Mesnil, 1952.—Afrotropical: D.R. Congo, South Africa.
Nemorilloides carbonata Mesnil, 1952a: 10. Holotype male (MRAC). Type locality: D.R. Congo, Nord-Kivu, Rutshuru, 1285m.
flaviventris Brauer & Bergenstamm, 1891.—Afrotropical: South Africa.
Nemorilloides flaviventris Brauer & Bergenstamm, 1891: 356 [also 1891: 52]. Lectotype female (NHMW, not located by JEOH), by fixation of Townsend (1941: 111) (mention of “Ht female” from Cape of Good Hope in NHMW is regarded as a lectotype fixation). Type locality: South Africa, Western Cape, Cape of Good Hope [as “Cap b. sp.” = “Cap Bonae Spei”].
Genus PARATRYPHERA Brauer & Bergenstamm, 1891
PARATRYPHERA Brauer & Bergenstamm, 1891: 328 [also 1891: 24]. Type species: Paratryphera handlirschii Brauer & Bergenstamm, 1891 (= Chetina palpalis Rondani, 1859), by monotypy [Palaearctic].
sordida (Villeneuve, 1916).—Afrotropical: Botswana, Kenya, South Africa, Tanzania, Uganda, Yemen.
Zenillia sordida Villeneuve, 1916c: 485. Holotype male (SAMC, not located by JEOH). Type locality: South Africa, KwaZulu-Natal, Durban.
sordia. Incorrect subsequent spelling of sordida Villeneuve, 1916 (Curran 1927d: 333).
Note: Paratryphera sordida (Villeneuve, 1916) of current authors is likely a species complex but is treated here as a single species pending further study.
Possibly undescribed spp.: Kenya, South Africa (both records based on specimens in BMNH, Crosskey 1984: 270).
Genus PHOROCEROSOMA Townsend, 1927
PHOROCEROSOMA Townsend, 1927c: 61. Type species: Phorocerosoma forte Townsend, 1927 (= Masicera vicaria Walker, 1856), by original designation [Oriental].
aberrans Verbeke, 1962.—Afrotropical: Rwanda.
Phorocerosoma aberrans Verbeke, 1962a: 170. Holotype female (IRSNB). Type locality: Rwanda, near mouth of Sebeya River, Gisenyi [as “Kisenyi”].
Note: The name Phorocerosoma aberrans was made available by Verbeke (1962a: 170) in a key that was apparently intended to precede the full description by Verbeke (1962b: 32, as “Phorocerosoma aberrans n. sp.”). No specimens were mentioned in the first work but a single specimen, a female “Type”, was cited in the second. Since the nominal species was clearly based on a single specimen, we regard the “Type” as the holotype of P. aberrans by monotypy in Verbeke (1962a).
albifacies Verbeke, 1962.—Afrotropical: Cameroon, D.R. Congo.
Phorocerosoma albifacies Verbeke, 1962a: 170. Lectotype female (IRSNB), by fixation of Verbeke (1962b: 32) (examination of “Type, 1♀” from Beni is regarded as a lectotype fixation). Type locality: D.R. Congo, Nord-Kivu, Beni.
Note: The name Phorocerosoma albifacies was made available by Verbeke (1962a: 170) in a key that was apparently intended to precede the full description by Verbeke (1962b: 30, as “Phorocerosoma albifacies n. sp.”). No specimens were mentioned in the first work but two females, “Type” and “Paratype”, were cited in the second. We regard the “Type” as the lectotype of P. albifacies by fixation of Verbeke (1962b: 32).
caparti Verbeke, 1962.—Afrotropical: Burundi, D.R. Congo, Tanzania, Uganda.
Phorocerosoma caparti Verbeke, 1962a: 171. Lectotype male (IRSNB), by fixation of Verbeke (1962b: 24) (examination of “Type, 1♂” of Phorocerosoma vicina Verbeke from Rutshuru is regarded as a lectotype fixation for Phorocerosoma caparti Verbeke). Type locality: D.R. Congo, Nord-Kivu, Rutshuru.
Phorocerosoma vicina Verbeke, 1962b: 22 (junior objective synonym of Phorocerosoma caparti Verbeke, 1962; both names based on same name-bearing type). Holotype male (IRSNB). Type locality: D.R. Congo, Nord-Kivu, Rutshuru.
Note: The name Phorocerosoma caparti was made available by Verbeke (1962a: 171) in a key that was apparently intended to precede the full description by Verbeke (1962b: 22, as “Phorocerosoma vicina n. sp.”). No specimens were mentioned in the first work but a male “Type” and a series of “Paratypes” (of both sexes) were cited in the second. No explanation was given for proposing the name P. vicina in Verbeke (1962b) for what was named P. caparti in Verbeke (1962a). Verbeke (1962b) did not mention the name P. caparti and therefore we treat P. vicina as a separate nominal species (as did Crosskey 1980b: 862) rather than as a replacement name or incorrect subsequent spelling, with both P. caparti and P. vicina based on the same name-bearing type. We regard the “Type” (Verbeke 1962b: 24) as both the holotype of P. vicina and the lectotype of P. caparti (by fixation of Verbeke 1962b: 24), making the two names objective synonyms.
echinum Verbeke, 1962.—Afrotropical: D.R. Congo.
Phorocerosoma echina Verbeke, 1962a: 170. Holotype male (MRAC). Type locality: D.R. Congo, Katanga, Parc National de l’Upemba [as “P.N.U.”], Rivière Lufira, [subtributary] Rivière Senze, Kaziba [as “Kaziba, affl. g. Senze, s.-affl. dr. Lufira”], 1140m.
Note: The name Phorocerosoma echina was made available by Verbeke (1962a: 170) in a key that was apparently intended to precede the full description by Verbeke (1962b: 29, as “Phorocerosoma echina n. sp.”). No specimens were mentioned in the first work but a single specimen, a male “Type”, was cited in the second. Since the nominal species was clearly based on a single specimen, we regard the “Type” as the holotype of P. echina by monotypy in Verbeke (1962a).
elegans Verbeke, 1962.—Afrotropical: D.R. Congo.
Phorocerosoma elegans Verbeke, 1962a: 171. Holotype male (MRAC). Type locality: D.R. Congo, Orientale, Isiro [as “Paulis”].
Note: The name Phorocerosoma elegans was made available by Verbeke (1962a: 171) in a key that was apparently intended to precede the full description by Verbeke (1962b: 28, as “Phorocerosoma elegans n. sp.”). No specimens were mentioned in the first work but a single specimen, a male “Type”, was cited in the second. Since the nominal species was clearly based on a single specimen, we regard the “Type” as the holotype of P. elegans by monotypy in Verbeke (1962a).
forcipatum Verbeke, 1962.—Afrotropical: D.R. Congo.
Phorocerosoma forcipata Verbeke, 1962a: 171. Lectotype male (IRSNB), by fixation of Verbeke (1962b: 27) (examination of “Type, 1♂” from Rutshuru is regarded as a lectotype fixation). Type locality: D.R. Congo, Nord-Kivu, Rutshuru.
Note: The name Phorocerosoma forcipata was made available by Verbeke (1962a: 171) in a key that was apparently intended to precede the full description by Verbeke (1962b: 25, as “Phorocerosoma forcipata n. sp.”). No specimens were mentioned in the first work but a male and female, as “Type” and “Paratype” respectively, were cited in the second. We regard the “Type” as the lectotype of P. forcipata by fixation of Verbeke (1962b: 27).
pilipes (Villeneuve, 1916).—Afrotropical: D.R. Congo, Madagascar, Mauritius, Nigeria, Sierra Leone, South Africa, Uganda.
Exorista pilipes Villeneuve, 1916c: 483. Lectotype male (IRSNB), by designation of Verbeke (1962b: 21). Type locality: South Africa, KwaZulu-Natal, Durban.
postulans (Walker, 1861).—Misidentification, not Afrotropical [known from Palaearctic, Oriental and Australasian regions].
Note: An unknown species was recorded as Phorocerosoma anomala Baranov, 1936 [properly “anomalum” in this combination] (currently a synonym of Phorocerosoma postulans (Walker, 1861), see Crosskey 1966b: 108 and Sabrosky and Crosskey 1969: 49) from Kenya and Tanzania by Mesnil (1959: 4) and from “tropical Africa” by Crosskey (1973b: 144, 1976: 225). Misidentifications (not recorded from the Afrotropical Region by Crosskey 1980b, O’Hara et al. 2009: 87).
Genus ZELINDOPSIS Anonymous, 1946
Zelindopsis Villeneuve, 1943c: 101. Nomen nudum (proposed after 1930 without designation of type species from four included species) (see note and Evenhuis et al. 2008: 34).
Zelindopsis Anonymous in Imperial Institute of Entomology, 1946: 208. Type species: Zelindopsis duplaria Villeneuve, 1943, by monotypy (see Evenhuis et al. 2008: 34).
Note: Villeneuve (1943c: 100) treated three species of Zenillia Robineau-Desvoidy, 1830 as forming “un petit groupe homogène”, including new species Zenillia stativa. Villeneuve (1943c: 101) then described a fourth species, duplaria, placing it and the preceding three species in his new genus Zelindopsis. Zelindopsis Villeneuve, 1943 is a nomen nudum because it was proposed after 1930 without designation of a type species from four included species (not three included species as stated by Evenhuis et al. 2008: 34).
bicincta (Villeneuve, 1916).—Afrotropical: Ghana, Nigeria, South Africa, Tanzania.
Zenillia bicinta Villeneuve, 1916c: 487. Lectotype male (IRSNB), by fixation of Villeneuve (1943c: 101) (treatment of the single male syntype from Nigeria as the “type” is regarded as a lectotype fixation). Type locality: northern Nigeria.
Zenillia bicincta denudata Villeneuve, 1943c: 101. Holotype male (IRSNB) (this is also the single paralectotype of Zenillia bicincta Villeneuve, 1916). Type locality: Ghana, Aburi.
Zenillia bicincta aristata Villeneuve, 1943c: 101. Holotype male (IRSNB). Type locality: South Africa, “Colonie du Cap” (former Cape Province, corresponding to the present-day Western Cape, Eastern Cape, Northern Cape, and North West [in part] provinces).
bicinta. Incorrect original spelling of bicincta Villeneuve, 1916 (Villeneuve 1916c: 487).
Note: The specific epithet of Zenillia bicinctaVilleneuve (1916c: 487) was originally published as bicinta but subsequent authors (e.g., Villeneuve 1943c, Mesnil 1959, Verbeke 1962a, Crosskey 1980b) used the spelling bicincta. The spelling bicincta is an incorrect subsequent spelling according to Article 33.3 of the Code (ICZN 1999) but because it is in prevailing usage and is attributed to Villeneuve (1943c), it is deemed to be the correct original spelling in compliance with Article 33.3.1.
cornuta Verbeke, 1962.—Afrotropical: D.R. Congo.
Zelindopsis cornuta Verbeke, 1962a: 168, 169. Holotype male (IRSNB). Type locality: D.R. Congo, Nord-Kivu, Rutshuru.
duplaria Villeneuve, 1943.—Afrotropical: Tanzania.
Zelindopsis duplaria Villeneuve, 1943c: 101. Holotype male (not located). Type locality: Tanzania.
Note: Villeneuve (1943c: 101) described duplaria and placed it in Zelindopsis. He did not use the combination Zenillia duplaria (cf. Zenillia stativa Villeneuve, 1943) and hence duplaria is treated as described in Zelindopsis.
illita (Villeneuve, 1916).—Afrotropical: Burundi, South Africa, Tanzania, Uganda, Zimbabwe.
Zenillia illita Villeneuve, 1916c: 486 (as “Zenillia (Pales?) illita”). Holotype female (IRSNB). Type locality: South Africa, KwaZulu-Natal, Durban.
Note: Villeneuve (1916c) wrote in his description of Zenillia illita that the “type is a ♀” (p. 486) but further on wrote “Natal, 1♂, Durban, S. Afric. Museum [= SAMC]” (p. 487), evidently referring to the same specimen. Villeneuve (1943c: 100) later confirmed the sex of the holotype as female, writing “La femelle a seule été décrite”. Verbeke (1962b: 169) cited the type as a male from “Natal” in IRSNB. We confirm that the holotype is in IRSNB as stated by Verbeke, but is a female, not a male.
nigripalpis Verbeke, 1962.—Afrotropical: D.R. Congo.
Zelindopsis nigripalpis Verbeke, 1962a: 169. Holotype male (IRSNB). Type locality: D.R. Congo, Nord-Kivu, Rutshuru.
nigrocauda (Curran, 1927).—Afrotropical: D.R. Congo.
Phorocera nigrocauda Curran, 1927c: 10. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
nitidicauda (Curran, 1940).—Afrotropical: South Africa.
Phorocera nitidicauda Curran, 1940: 7. Holotype male (SANC). Type locality: South Africa, Gauteng, Pretoria.
Note: Phorocera nitidicauda Curran, 1940 is “almost certainly” a synonym of Zenillia illita Villeneuve, 1916 according to Crosskey (1984: 270).
nudapex (Curran, 1940).—Afrotropical: South Africa, Zimbabwe.
Phorocera nudapex Curran, 1940: 5. Holotype female (AMNH). Type locality: Zimbabwe, Nyanga [as “Inyanga”].
Note: Phorocera nudapex Curran, 1940 is “possibly” a synonym of Zenillia illita Villeneuve, 1916 according to Crosskey (1984: 270).
stativa (Villeneuve, 1943).—Afrotropical: D.R. Congo.
Zenillia stativa Villeneuve, 1943c: 101. Holotype male (IRSNB). Type locality: D.R. Congo, Nord-Kivu, Mukule, 1800m [ca. 1°20′S 29°15′E].
Note: In an unusual nomenclatural action, Villeneuve (1943c: 101) described his new species stativa in Zenillia Robineau-Desvoidy, 1830 but three paragraphs later placed it in his new genus Zelindopsis along with three other species. We interpret stativa as intentionally described in the combination Zenillia stativa and then moved to Zelindopsis, rather than described as Zelindopsis stativa.
villeneuvei Verbeke, 1962.—Afrotropical: D.R. Congo.
Zelindopsis villeneuvei Verbeke, 1962a: 168, 169. Holotype male (IRSNB). Type locality: D.R. Congo, northwest of Lake Tanganyika [as “N.W. Tanganika”] (not Tanzania as cited by Crosskey 1980b: 863, see note).
Note: The holotype of Zelindopsis villeneuvei Verbeke, 1962 was collected by [Rudolf] Grauer in 1910 from “N.W. Tanganika” according to both Verbeke (1962a: 169) and label data (holotype examined by PC). Grauer collected a wide variety of animals including insects, snakes, birds and mammals from “N.W. Tanganika” in 1910, as evidenced by numerous works citing his specimens. A few sources have interpreted “N.W. Tanganika” as Northwest Tanzania (including Crosskey 1980b: 863) but most have treated it as northwest (or northwest shore) of Lake Tanganyika; i.e., in D.R. Congo. One seemingly authoritative reference with the latter interpretation is Chapin (1928: 7, 8), and we accept this view.
ugandana (Curran, 1940).—Afrotropical: Uganda.
Phorocera ugandana Curran, 1940: 3. Holotype male (BMNH). Type locality: Uganda, Lake Kibivera [not located].
zenia (Curran, 1940).—Afrotropical: Uganda.
Phorocera zenia Curran, 1940: 10. Holotype male (BMNH). Type locality: Uganda, Kampala.
Tribe GONIINI
Genus AGAEDIOXENIS Villeneuve, 1939
GAEDIOXENIS Villeneuve, 1937: 206. Nomen nudum (proposed after 1930 without designation of type species from two included species).
GAEDIOXENIS Villeneuve, 1939: 1. Nomen nudum (proposed after 1930 without designation of type species from two included species).
AGAEDIOXENIS Villeneuve, 1939: 2 (as subgenus of Gaedioxenis Villeneuve, 1937 [nomen nudum]). Type species: Gaedioxenis (Agaedioxenis) brevicornis Villeneuve, 1939, by monotypy
GAEDIOXENIS Townsend, 1943a: 335. Type species: Gaedioxenis setifrons Villeneuve, 1937, by original designation.
Note: Villeneuve (1937) described the genus Gaedioxenis with two new species, then two years later Villeneuve (1939) added another new species to the genus and created for it the new subgenus Agaedioxenis. The name Gaedioxenis was a nomen nudum in both publications but Agaedioxenis was validly proposed in the second. The valid name of the genus is thus Agaedioxenis, as explained in more detail by Cerretti et al. (2015: 502) in their revision of Agaedioxenis Villeneuve, 1939 and Eugaedioxenis Cerretti, O’Hara & Stireman, 2015.
brevicornis (Villeneuve, 1939).—Afrotropical: South Africa, Zimbabwe.
Gaedioxenis (Agaedioxenis) brevicornis Villeneuve, 1939: 1. Holotype male (BMNH). Type locality: Zimbabwe, Mutare [as “Umtali”] District.
Gaedioxenis propinqua Villeneuve, 1939: 2. Holotype female (not located). Type locality: South Africa, KwaZulu-Natal.
Note: The relative priority of Gaedioxenis (Agaedioxenis) brevicornis Villeneuve, 1939 and Gaedioxenis propinqua Villeneuve, 1939, when the two are treated as synonyms, was established by Cerretti et al. (2015: 506), as the First Reviser (Article 24.2.2 of the Code, ICZN 1999).
kirkspriggsi Cerretti, O’Hara & Stireman, 2015.—Afrotropical: South Africa.
Agaedioxenis kirkspriggsi Cerretti, O’Hara & Stireman inCerretti et al., 2015: 507. Holotype male (NMB). Type locality: South Africa, Free State, Harrismith, Mooihekkop, ca. 1800m (28°10′50.0″S 29°10′51.1″E).
setifrons (Villeneuve, 1937).—Afrotropical: South Africa.
Gaedioxenis setifrons Villeneuve, 1937a: 207. Holotype female (CNC). Type locality: South Africa, Western Cape, Stellenbosch.
succulentus Cerretti, O’Hara & Stireman, 2015.—Afrotropical: South Africa.
Agaedioxenis succulentus Cerretti, O’Hara & Stireman inCerretti et al., 2015: 507. Holotype male (MZUR). Type locality: South Africa, Western Cape, Ceres Bergfynbos Reserve, 459m (33°23′1.91″S 19°17′20.16″E).
timidus Cerretti, O’Hara & Stireman, 2015.—Afrotropical: South Africa.
Agaedioxenis timidus Cerretti, O’Hara & Stireman inCerretti et al., 2015: 508. Holotype male (CNC). Type locality: South Africa, Western Cape, Cape Town.
Genus BLEPHARELLA Macquart, 1851
BLEPHARELLA Macquart, 1851b: 176 [also 1851b: 203]. Type species: Blepharella lateralis Macquart, 1851, by original designation [Oriental].
PODOMYIA Brauer & Bergenstamm, 1889: 96 [also 1890: 28]. Type species: Eurigaster setosa Doleschall, 1858 (= Blepharella lateralis Macquart, 1851), by monotypy [Oriental].
CONGOCHRYSOSOMA Townsend, 1916a: 174. Type species: Congochrysosoma snyderi Townsend, 1916, by original designation.
PHRYXOSTURMIA Townsend, 1927c: 68. Type species: Phryxosturmia jacobsoni Townsend, 1927 (= Blepharella lateralis Macquart, 1851), by original designation [Oriental].
AFROSTURMIA Curran, 1927f: 126. Type species: Afrosturmia orbitalis Curran, 1927, by original designation. Syn. n.
APILIA Malloch, 1930a: 345. Type species: Apilia cilifera Malloch, 1930 [= Blepharella lateralis Macquart, 1851], by original designation [Australasian].
PUJOLINA Mesnil, 1968b: 2. Type species: Pujolina bicolor Mesnil, 1968, by original designation.
Note: Macquart (1851b: 177 [also 1851b: 204]) noted, about his new genus Blepharella, “Le type est asiatique”. This statement is accepted as a type species designation for Blepharella of the single included species from India, Blepharella lateralis Macquart.
Afrosturmia Curran, 1927 was treated as a monotypic genus by Crosskey (1980b: 867) but is here placed in synonymy with Blepharella Macquart, 1851.
abana (Curran, 1927).—Afrotropical: Angola, Tanzania.
Sturmia abana Curran, 1927f: 122. Holotype male (BMNH). Type locality: Tanzania, Morogoro.
alacris (Curran, 1927).—Afrotropical: Malawi, Nigeria, Tanzania.
Sturmia alacris Curran, 1927f: 123. Holotype male (BMNH). Type locality: Tanzania, Morogoro.
analis (Curran, 1927).—Afrotropical: D.R. Congo, Kenya, Somalia, Tanzania, Zimbabwe.
Sturmia analis Curran, 1927f: 120. Holotype male (BMNH). Type locality: Kenya, Narok [as “Narok, Masai Reserve”, ca. 1°5′S 35°52′E].
arrogans (Curran, 1927).—Afrotropical: D.R. Congo.
Sturmia arrogans Curran, 1927c: 16. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
atricauda Mesnil, 1970.—Afrotropical: Zimbabwe.
Blepharella (Congochrysosoma) atricauda Mesnil, 1970b: 97. Holotype male (CNC). Type locality: Zimbabwe, Hurungwe [as “Urungwe”], Gota Gota.
aurifrons (Villeneuve, 1916).—Afrotropical: D.R. Congo, Kenya, Malawi, Sierra Leone, South Africa, Tanzania, Uganda.
Sturmia (Crossocosmia) aurifrons Villeneuve, 1916c: 475. Syntypes, males and females (SAMC [1 male examined by JEOH], other unspecified collections [as “etc.”]). Type localities: Malawi (Mt. Mulanje [as “Mt. Mlanje”]), Sierra Leone, South Africa (KwaZulu-Natal, Mfongosi [as “M’fongosi, Zululand”, ca. 28°43′S 30°49′E]), and Uganda.
bicolor (Mesnil, 1968).—Afrotropical: C.A. Republic, D.R. Congo.
Pujolina bicolor Mesnil, 1968b: 3. Holotype female (MNHN). Type locality: C.A. Republic.
carbonata Mesnil, 1952.—Afrotropical: D.R. Congo.
Blepharella (Blepharella) setigera carbonata Mesnil, 1952c: 235. Holotype, unspecified sex [male, see Cooper and O’Hara 1996: 18] (CNC). Type locality: D.R. Congo, Nord-Kivu, Kabasha [Escarpment], “Chambi” [probably Tshambi, ca. 0°44′S 29°13′E].
chionaspis (Bezzi, 1908).—Afrotropical: D.R. Congo.
Winthemia chionaspis Bezzi, 1908c: 382. Holotype male (?IRSNB). Type locality: D.R. Congo, Orientale [as “Haut-Congo”].
confusa Mesnil, 1952.—Afrotropical: South Africa.
Blepharella (Blepharella) setigera confusa Mesnil, 1952c: 235. Holotype male (CNC). Type locality: South Africa.
Note: Mesnil (1952c: 235) gave the type locality of Blepharella setigera confusa as South Africa. The holotype has a label with a place name but the name is partly obscured by a spot of black ink and is unreadable.
erebiae Mesnil, 1970.—Afrotropical: Malawi.
Blepharella (Congochrysosoma) erebiae Mesnil, 1970b: 96. Holotype female (CNC). Type locality: Malawi, Mt. Mulanje [as “Mt. Mlanje”].
fallaciosa Mesnil, 1970.—Afrotropical: Uganda.
Blepharella (Congochrysosoma) fallaciosa Mesnil, 1970b: 96. Holotype male (CNC). Type locality: Uganda, Entebbe.
fascipes (Villeneuve, 1943).—Afrotropical: D.R. Congo, Ethiopia, South Africa.
Sturmia fascipes Villeneuve, 1943a: 37. Holotype male (CNC). Type locality: D.R. Congo, Katanga, Sankisia.
fuscicosta (Curran, 1927).—Afrotropical: D.R. Congo, Ghana, Guinea, Malawi, Uganda.
Sturmia (Crossocosmia) fuscicosta Curran, 1927a: 10. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
fuscipennis Mesnil, 1952.—Afrotropical: D.R. Congo.
Blepharella (Blepharella) fuscipennis Mesnil, 1952c: 235. Holotype male (CNC). Type locality: D.R. Congo, Orientale, Penghe [near Mambasa, 1°22′24″N 29°4′34″E].
grandis (Curran, 1927).—Afrotropical: D.R. Congo.
Sturmia grandis Curran, 1927a: 13. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
haemorrhoa Mesnil, 1970.—Afrotropical: Madagascar.
Blepharella (Congochrysosoma) haemorrhoa Mesnil, 1970b: 95. Holotype male (MNHN). Type locality: Madagascar, Toliara, Andronobe.
hova Mesnil, 1952.—Afrotropical: Madagascar, South Africa.
Blepharella (Blepharella) hova Mesnil, 1952c: 235. Holotype male (MNHN). Type locality: Madagascar, “Merinon” [not located].
imitator (Curran, 1927).—Afrotropical: D.R. Congo, Uganda.
Sturmia imitator Curran, 1927a: 13. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
instabilis (Curran, 1927).—Afrotropical: Malawi, South Africa.
Sturmia instabilis Curran, 1927f: 124. Holotype male (SANC). Type locality: South Africa, KwaZulu-Natal, Port Shepstone.
intensica (Curran, 1927).—Afrotropical: D.R. Congo.
Sturmia (Crossocosmia) intensica Curran, 1927a: 17. Lectotype male (AMNH), by designation of Arnaud (1963: 129). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
laetabilis (Curran, 1927).—Afrotropical: D.R. Congo, Ghana, Nigeria, Sierra Leone.
Sturmia laetabilis Curran, 1927f: 112, 114. Syntypes, 3 males and 2 females (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
Note: Authorship of Sturmia laetabilis was attributed to Villeneuve (1933: 279) by Crosskey (1980b: 868). However, the characters given for S. laetabilis by Curran (1927f) in his key to the African species of Sturmia (pp. 112 [male], 114 [female]) validate the name from this work. Curran (1927f: 126) cited the author as “Villeneuve (in litt.?)”, but the descriptive details given for this species were his own: “I have several specimens from the Belgian Congo determined by Villeneuve, but have seen no description”. Curran (1928b) keyed S. laetabilis (pp. 389, 391) and treated it in his text (p. 394), again with Villeneuve as author, and cited 3 males and 2 females from “Stanleyville”. These specimens are assumed to be the same as those examined by Curran (1927f) and thus are accepted as the original syntypes of S. laetabilis. Villeneuve (1933: 279) gave characters to separate “Sturmia rubricosa n. sp.” and “Sturmia laetabilis” but interestingly did not name the latter as a new species nor cite it with an author’s name. No locality data was provided by Villeneuve for S. laetabilis. The distribution of Blepharella laetabilis (Curran) is recognized here as the type locality in D.R. Congo and the three countries listed for the species by Crosskey (1980b): Ghana, Nigeria, and Sierra Leone.
lodosi Mesnil, 1968.—Afrotropical: Ghana.
Blepharella (Congochrysosoma) lodosi Mesnil, 1968b: 1. Holotype male (CNC). Type locality: Ghana, Tafo [suburb of Kumasi].
melita (Curran, 1927).—Afrotropical: D.R. Congo.
Sturmia melita Curran, 1927c: 12. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
neglecta Mesnil, 1968.—Afrotropical: D.R. Congo.
Blepharella (Congochrysosoma) neglecta Mesnil, 1968b: 2. Holotype male (CNC). Type locality: D.R. Congo, Nord-Kivu, Walikale [ca. 1°25′S 28°00′E].
oldi Mesnil, 1952.—Afrotropical: Malawi.
Blepharella (Blepharella) oldi Mesnil, 1952c: 235. Holotype male (CNC). Type locality: Malawi, “Ruo” (“Aluona Ruo Dist” according to label data, Cooper and O’Hara 1996: 18; likely somewhere in the Shire Valley of the former Ruo District, ca. 17°S 35°E [given as Ruo, Tanzania by O’Hara 1996: 150 and Cooper and O’Hara 1996: 18, in error]).
orbitalis (Curran, 1927).—Afrotropical: Ghana. Comb. n.
Afrosturmia orbitalis Curran, 1927f: 127. Holotype male (BMNH). Type locality: Ghana, Ashanti.
Note: Afrosturmia orbitalis Curran, 1927 was treated as the sole species of Afrosturmia by Crosskey (1980b: 867) but is moved here to Blepharella Macquart, 1851.
pellucida Mesnil, 1970.—Afrotropical: D.R. Congo.
Blepharella (Congochrysosoma) pellucida Mesnil, 1970b: 98. Holotype male (CNC). Type locality: D.R. Congo, Équateur, Lulonga [ca. 0°37′N 18°22′E].
perfida Mesnil, 1970.—Afrotropical: D.R. Congo.
Blepharella (Congochrysosoma) perfida Mesnil, 1970b: 96. Holotype male (CNC). Type locality: D.R. Congo, Katanga, Kafakumba [ca. 9°41′N 23°46′E].
picturata (Curran, 1927).—Afrotropical: Kenya, Uganda.
Sturmia picturata Curran, 1927f: 122. Holotype female (BMNH). Type locality: Uganda, “Kukedi” [not located].
rex (Curran, 1927).—Afrotropical: D.R. Congo, Tanzania, Uganda.
Sturmia rex Curran, 1927a: 14. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
rubricosa (Villeneuve, 1933).—Afrotropical: Malawi.
Sturmia rubricosa Villeneuve, 1933: 279. Syntypes, males (“plusieurs”) and 1 female (not located). Type locality: Malawi.
ruficauda Mesnil, 1952.—Afrotropical: South Africa.
Blepharella (Blepharella) setigera ruficauda Mesnil, 1952c: 235. Holotype male (CNC). Type locality: South Africa, Gauteng, Sydenham.
setifacies (Curran, 1927).—Afrotropical: D.R. Congo, Uganda.
Sturmia setifacies Curran, 1927c: 12. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
Sturmia femineum Curran, 1927c: 14. Holotype female (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
Winthemia orbitalis Villeneuve, 1934d: 68 (junior secondary homonym of Afrosturmia orbitalis Curran, 1927). Holotype male (CNC). Type locality: D.R. Congo, Nord-Kivu, “Moko Lesse” (“Moko” is “Moho” on the locality label of the holotype, Cooper and O’Hara 1996: 77) [Lesse at ca. 0°45′N 29°46′E, Moho (or Moko) is presumed to be nearby].
Blepharella (Congochrysosoma) erronea Mesnil, 1970b: 95 (nomen novum for Winthemia orbitalis Villeneuve, 1934).
Note: The relative priority of Sturmia setifacies Curran, 1927 and Sturmia femineum Curran, 1927, when the two are treated as synonyms, was established by Crosskey (1980b: 868), as the First Reviser (Article 24.2.2 of the Code, ICZN 1999).
setigera (Corti, 1895).—Afrotropical: “widespread Afrotrop. Reg.” (Crosskey 1980b: 869), including D.R. Congo, Ethiopia, Kenya, Malawi, Nigeria, Sierra Leone, Uganda. Palaearctic: M. East (M. East [Iran, Zeegers and Majnon Jahromi 2015: 539]).
Podomyia setigera Corti, 1895: 135. Type(s), male (?MCSN). Type locality: Ethiopia, Jubba River, “Arussi Galla, Ganale Guddà” [most likely a valley of the upper Ganale River, a tributary of the Jubba River on the eastern edge of the Arussi and Bale Mountains, ca. 7°0′N 40°30′E].
setigena. Incorrect subsequent spelling of setigera Corti, 1895 (Zeegers and Majnon Jahromi 2015: 540, etc.).
seydeli (Mesnil, 1949).—Afrotropical: D.R. Congo.
Zygobothria seydeli Mesnil, 1949a: 92. Holotype male (MRAC). Type locality: D.R. Congo, Katanga, Lubumbashi [as “Elisabethville”].
snyderi (Townsend, 1916).—Afrotropical: D.R. Congo, Ghana, Guinea, Kenya, Malawi, Nigeria, Tanzania, Uganda.
Congochrysosoma snyderi Townsend, 1916a: 174. Holotype female (USNM). Type locality: D.R. Congo, Kasai-Occidental, Luebo.
Sturmia currani Villeneuve, 1933: 279 (named for Sturmia versatilis of Curran, 1927f, 1928b, not Villeneuve, 1910a). Syntypes, females (“plusieurs”) (not located). Type localities: D.R. Congo and Malawi.
Sturmia versatilis of Curran (1927f: 125, 1928b: 394), not Villeneuve, 1910. Misidentification (Villeneuve 1933: 279).
vasta (Karsch, 1886).—Afrotropical: Angola, Uganda.
Tachina vasta Karsch, 1886b: 341. Holotype, unspecified sex [female, examined by JEOH] (ZMHB). Type locality: Angola, Pungo Andongo.
versatilis (Villeneuve, 1910).—Afrotropical: D.R. Congo, Malawi, Nigeria, Sudan. Palaearctic: ?N. Africa (NE. Africa) (see note).
Sturmia versatilis Villeneuve, 1910a: 253. Type(s), male (1 male in CNC). Type locality: D.R. Congo (as “Congo”, p. 249).
versatalis. Incorrect subsequent spelling of versatilis Villeneuve, 1910 (Curran 1928b: 389).
Note: Villeneuve (1910a) described four species from “Congo”. Curran (1927f: 122) treated one of them (Sturmia aureiventris Villeneuve, 1910) as described from D.R. Congo (as “Belgian Congo”), and used “Belgian Congo” and “Congo” interchangeably in this work and some others. We think it likely that Villeneuve (1910a), like Curran, used “Congo” in the sense of present-day D.R. Congo. However, Crosskey (1980b) interpreted Villeneuve’s Congo as the present-day country of Congo. Villeneuve’s (1913c: 29) record of Sturmia versatilis Villeneuve, 1910 from Egypt, based on a male in BMNH, needs confirmation.
vivax (Curran, 1927).—Afrotropical: D.R. Congo, Nigeria.
Sturmia vivax Curran, 1927a: 15. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
vulnerata (Curran, 1927).—Afrotropical: D.R. Congo.
Sturmia vulnerata Curran, 1927c: 13. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
xanthaspis Mesnil, 1970.—Afrotropical: South Africa.
Blepharella (Congochrysosoma) xanthaspis Mesnil, 1970b: 97. Holotype male (CNC). Type locality: South Africa, KwaZulu-Natal, Eshowe [as “Eshova”, misprint].
Genus BLEPHARELLINA Mesnil, 1952
BLEPHARELLINA Mesnil, 1949b: 104. Nomen nudum (proposed after 1930 without designation of type species; no included species) (see Evenhuis and O’Hara 2008: 65).
BLEPHARELLINA Mesnil, 1950c: 105 (as subgenus of Blepharella Macquart, 1851). Nomen nudum (proposed after 1930 without designation of type species; no included species) (see Evenhuis and O’Hara 2008: 65).
BLEPHARELLINA Mesnil, 1952c: 234 (as subgenus of Blepharella Macquart, 1851). Type species: Blepharella (Blepharellina) picta Mesnil, 1952, by monotypy (see Evenhuis and O’Hara 2008: 65).
picta (Mesnil, 1952).—Afrotropical: Nigeria.
Blepharella (Blepharellina) picta Mesnil, 1952: 234. Holotype, unspecified sex [female, see Cooper and O’Hara 1996: 18] (CNC). Type locality: Nigeria, Oshogbo.
Genus BLEPHARIPA Rondani, 1856
BLEPHARIPA Rondani, 1856: 71. Type species: Erycia ciliata Macquart, 1834 (as “Masicera ciliata Macq.”) (= Tachina pratensis Meigen, 1824), by original designation.
BLEPHARIPODA Brauer & Bergenstamm, 1889: 96 [also 1890: 28] (junior homonym of Blepharipoda Randall, 1840). Type species: Nemoraea scutellata Robineau-Desvoidy, 1830 (= Tachina pratensis Meigen, 1824), by monotypy.
pratensis (Meigen, 1824).—Misidentification, not Afrotropical [known from Palaearctic Region].
Note: An unknown species was recorded as “Sturmia scutellata, Desvoidy” (originally described as Nemoraea scutellata Robineau-Desvoidy, 1830, currently a synonym of Tachina pratensis Meigen, 1824) from Uganda by Curran (1927f: 123). Misidentification (not recorded from the Afrotropical Region by Crosskey 1980b, Herting and Dely-Draskovits 1993: 249).
Genus BRACHYCHAETOIDES Mesnil, 1970
BRACHYCHAETOIDES Mesnil, 1970b: 109 (as subgenus of Chlorolydella Townsend, 1933). Type species: Chlorolydella (Brachychaetoides) varipes Mesnil, 1970 (= Archiclops africanum Mesnil, 1968), by original designation.
Note: Brachychaetoides Mesnil, 1970 was treated as a synonym of Chlorolydella Townsend, 1933 by Crosskey (1980b: 877). It was later recognized as a genus by Crosskey (1984: 201, 295) with single species Brachychaetoides africanum (Mesnil, 1968).
africanum (Mesnil, 1968).—Afrotropical: Tanzania.
Archiclops africanum Mesnil, 1968b: 6. Holotype male (SMNS). Type locality: Tanzania, southwest side of Mt. Kilimanjaro [as “Kilimandjaro”], 3500m.
Chlorolydella (Brachychaetoides) varipes Mesnil, 1970b: 109. Holotype male (MNHN). Type locality: Tanzania, Mt. Kilimanjaro [as “Kilimandjaro”], 2800–3000m.
Note: Chlorolydella varipes Mesnil, 1970 was synonymized with Archiclops africanum Mesnil, 1968 by Crosskey (1984: 201, 295). Archiclops africanum was earlier treated as a species of Gymnophryxe Villeneuve, 1922 by Crosskey (1980b: 878).
violacea (Curran, 1927).—Afrotropical: Kenya. Comb. n.
Campylochaeta violacea Curran, 1927d: 337. Holotype male (BMNH). Type locality: Kenya, Kabete [ca. 1°16′S 36°43′E, near Nairobi].
Note: Campylochaeta violacea Curran, 1927 was treated as a species of Chlorolydella Townsend, 1933 by Crosskey (1980b: 877, 1984: 286) but is moved here to Brachychaetoides Mesnil, 1970.
Undescribed spp.: Kenya (TAU), Malawi (TAU), South Africa (MZUR, NMB) (examined by PC).
Genus CADURCIA Villeneuve, 1926
CADURCIA Villeneuve, 1926c: 243. Type species: Masicera casta Rondani, 1861, by subsequent designation of Townsend (1936b: 256) [Palaearctic].
ARGYROPHYLACOIDES Townsend, 1933: 477. Type species: Degeeria zetterstedtii Karsch, 1886, by original designation.
auratocauda (Curran, 1934).—Afrotropical: Côte d’Ivoire, D.R. Congo, Ghana, Nigeria, Sierra Leone.
Sturmia auratocauda Curran, 1934b: 2. Holotype male (BMNH). Type locality: Nigeria, Ibadan.
borbonensis Villeneuve, 1926.—Afrotropical: Réunion.
Cadurcia borbonensis Villeneuve, 1926c: 245. Syntypes, 4 males (1 male in CNC, 1 male in NHMW). Type localities: Réunion and “un ♂ de la collection Strobl (d’où?)”.
depressa Villeneuve, 1926.—Afrotropical: D.R. Congo.
Cadurcia depressa Villeneuve, 1926c: 244. Syntypes, 2 males (1 male in CNC). Type locality: D.R. Congo, Katanga, Kayombo.
fascicauda (Curran, 1934).—Afrotropical: South Africa.
Sturmia fascicauda Curran, 1934b: 3. Holotype male (SANC). Type locality: South Africa, Eastern Cape, East London.
lucens Villeneuve, 1926.—Afrotropical: Malawi, Mauritius, Nigeria, South Africa, Uganda.
Cadurcia lucens Villeneuve, 1926c: 244. Lectotype male (BMNH), by designation of Crosskey (1976: 265). Type locality: Nigeria, Ilorin.
Masicera casta of authors (e.g., Verbeke 1970: 272, as “Cadurcia casta”; Crosskey 1980b: 860, under “Cadurcia lucens”), not Rondani, 1861. Misidentification (see note).
Note: Crosskey (1980b: 869) included Cadurcia vanderwulpi Baranov, 1938 (described from India) in synonymy with Cadurcia lucens Villeneuve, 1926, but we have followed Herting and Dely-Draskovits (1993: 245) in treating the former as a valid name. Similarly, we have followed Herting and Dely-Draskovits (1993: 243) in treating C. casta (Rondani, 1861) as a strictly Palaearctic species and not as a questionable synonym of C. lucens as listed by Crosskey (1980b: 869). If C. casta and C. lucens are conspecific then the former name has priority.
mesnili Verbeke, 1962.—Afrotropical: D.R. Congo.
Cadurcia mesnili Verbeke, 1962b: 53. Holotype male (IRSNB). Type locality: D.R. Congo, Nord-Kivu, Parc National des Virunga [as “P.N.A”, former Parc National Albert], Goma-Sake route, Buheno.
plutellae van Emden, 1942.—Afrotropical: Kenya.
Cadurcia plutellae van Emden, 1942: 223. Holotype male (BMNH). Type locality: Kenya, Nairobi.
semiviolacea Villeneuve, 1926.—Afrotropical: South Africa.
Cadurcia semiviolacea Villeneuve, 1926c: 245 (as “semiviolacea (B. B. i. litt.)”). Syntypes, 2 females (NHMW). Type locality: South Africa, Western Cape, Cape of Good Hope [as “Cap”].
Note: Villeneuve (1926c: 245) wrote that Cadurcia semiviolacea was “Représenté par 2 ♀♀ dans la collection v. Winthem du Muséum de Vienne”. Villeneuve began the next paragraph with “Ces deux ♂♂”, which was a lapsus for two females. The two female syntypes in NHMW were examined by JEOH.
versicauda (Curran, 1934).—Afrotropical: Angola, South Africa, Tanzania.
Sturmia versicauda Curran, 1934b: 4. Holotype male (BMNH). Type locality: South Africa, KwaZulu-Natal, Weenen [as “Wernen”, ca. 28°51′S 30°4′E].
vinsoni Mesnil, 1952.—Afrotropical: Mauritius.
Cadurcia vinsoni Mesnil, 1952c: 214. Holotype, unspecified sex (BMNH). Type locality: Mauritius, Chebel.
zetterstedtii (Karsch, 1886).—Afrotropical: Angola, Congo, Guinea, Nigeria, Senegal, Yemen.
Degeeria zetterstedtii Karsch, 1886b: 342. Holotype, unspecified sex [female, examined by JEOH] (ZMHB). Type locality: Angola, Pungo Andongo.
Sturmia albicauda Curran, 1934b: 3. Holotype male (AMNH). Type locality: Congo, “on board ship off Loango”.
albocauda. Incorrect original spelling of albicauda Curran, 1934 (Curran 1934b: 1).
Note: There are two original spellings for Sturmia albicauda in Curran (1934b): albocauda in the key (p. 1) and albicauda in the species header (p. 3). The correct original spelling was selected as albicauda by Crosskey (1980b: 869), as the First Reviser (Article 24.2.3 of the Code, ICZN 1999).
Possibly undescribed spp.: Yemen, as “Cadurcia sp. 1 cf. fascicauda” and “Cadurcia sp. 2” (Zeegers 2007: 378).
Genus CHAETOSTURMIA Villeneuve, 1915
CHAETOSTURMIA Villeneuve, 1915b: 193. Type species: Chaetosturmia barbata Villeneuve, 1915, by monotypy.
barbata Villeneuve, 1915.—Afrotropical: Madagascar.
Chaetosturmia barbata Villeneuve, 1915b: 194. Holotype male (NHMW). Type locality: Madagascar.
Genus CHLOROLYDELLA Townsend, 1933
CHLOROLYDELLA Townsend, 1933: 473. Type species: Chlorolydella caffrariae Townsend, 1933, by original designation.
CHLOROPHRYNO Townsend, 1933: 478. Type species: Gymnochaeta glauca Karsch, 1886 (as “Gymnocheta glauca”), by original designation.
Note: The relative priority of Chlorolydella Townsend, 1933 and Chlorophryno Townsend, 1933, when the two are treated as synonyms, was established by Mesnil (1954b: 347), as the First Reviser (Article 24.2.2 of the Code, ICZN 1999). Mesnil did not mention Chlorophryno but effectively synonymized it with Chlorolydella by placing its type species, Gymnochaeta glauca Karsch, 1886, in Chlorolydella.
bequaerti (Curran, 1940).—Afrotropical: Uganda.
Phorocera bequaerti Curran, 1940: 6. Holotype female (AMNH). Type locality: Uganda, Behungi [as “Behunge” in error, Arnaud 1963: 124, ca. 1°17′S 29°48′E].
caffrariae Townsend, 1933.—Afrotropical: South Africa, Tanzania, Uganda, Zimbabwe.
Chlorolydella caffrariae Townsend, 1933: 474. Holotype male (NHRS). Type locality: South Africa, “Caffraria” (also known as “Kaffraria”, a former region in Eastern Cape).
? Stomatomya metallica Villeneuve, 1916c: 475 (junior secondary homonym of Campylochaeta metallica Bezzi, 1908 and Phorocera metallica Becker, 1909). Syntypes, unspecified number and including at least 1 male (CNC, SAMC [not located by JEOH]). Type localities: South Africa (KwaZulu-Natal, Durban; KwaZulu-Natal, Mfongosi [as “M’fongosi, Zululand”]) and Zimbabwe (Harare [as “Salisbury”]).
glauca (Karsch, 1886).—Afrotropical: Angola, Burundi, Eritrea, Kenya, South Africa (new record, NMDA [PC]), Tanzania, Uganda.
Gymnochaeta glauca Karsch, 1886b: 339. Syntypes, two specimens of unspecified sex [females, examined by JEOH] (ZMHB). Type locality: Angola, Pungo Andongo (not “West Tanganyika” [i.e., Tanzania] as cited by Townsend 1933: 478, in error).
Campylochaeta metallica Bezzi, 1908b: 57. Holotype male (not located, not among the labelled types of Bezzi in MSNM examined by Arnaud 1982). Type locality: Eritrea, near Adi Keyh [also as Adi Kaie and other spellings, published as “Adi Caiè”, ca. 14°51′N 39°22′E].
Note: Townsend (1933: 478) mentioned the “Female holotype” of Gymnochaeta glauca Karsch, 1886 in ZMHB but did not restrict the term holotype to one of the two females in the type series and hence did not fix a lectotype.
metallica (Becker, 1909).—Afrotropical: Kenya.
Phorocera metallica Becker, 1909a: 117 (junior secondary homonym of Campylochaeta metallica Bezzi, 1908; not renamed while Campylochaeta metallica is in synonymy with Chlorolydella glauca (Karsch, 1886)). Holotype female (MNHN). Type locality: Kenya [as “Afrique orientale anglaise; Escarpment”, interpreted as Kenya by Crosskey 1980b: 877].
Note: The description of Phorocera metallica Becker, 1909 was repeated in Becker (1910a: 26) under the heading “Phorocera metallica, n. sp. ♀”.
pallidipes (Curran, 1927).—Afrotropical: Kenya.
Campylochaeta pallidipes Curran, 1927d: 338. Holotype male (not located). Type locality: Kenya, Kabete [ca. 1°16′S 36°43′E, near Nairobi].
schistacea Mesnil, 1955.—Afrotropical: Rwanda, South Africa.
Chlorolydella schistacea Mesnil, 1955: 365. Holotype, unspecified sex [male, see Cooper and O’Hara 1996: 24] (CNC). Type locality: Rwanda, Volcan Visoke [also known as Bisoke; published as “Bishoke”], Kibga, 2400m [ca. 1°29′S 29°31′E].
trochanterata (Villeneuve, 1934).—Afrotropical: South Africa.
Pales trochanterata Villeneuve, 1934c: 408. Syntypes, 2 males (not located). Type locality: South Africa.
venusta (Curran, 1928).—Afrotropical: Burundi, Kenya, Tanzania, Uganda.
Phorocera venusta Curran, 1928a: 238. Holotype male (BMNH). Type locality: Uganda, Rwenzori Range [as “Mount Ruwenzori”].
Genus DOLICHOCOLON Brauer & Bergenstamm, 1889
DOLICHOCOLON Brauer & Bergenstamm, 1889: 100 [also 1890: 32]. Type species: Dolichocolon paradoxum Brauer & Bergenstamm, 1889, by monotypy.
Note: A world revision of Dolichocolon Brauer & Bergenstamm, 1889 was published by Cerretti and Shima (2011).
africanum Mesnil, 1968.—Afrotropical: D.R. Congo, South Africa, Tanzania.
Dolichocolon africanum Mesnil, 1968c: 176. Holotype male (CNC). Type locality: D.R. Congo, Nord-Kivu, Rwindi, 1000m [ca. 0°47′S 29°17′E].
basilewskyi Cerretti & Shima, 2011.—Afrotropical: Uganda.
Dolichocolon basilewskyi Cerretti & Shima, 2011: 557. Holotype male (MRAC). Type locality: Uganda, Bugiri, 1400m (1°04′N 33°43′E).
bequaerti Cerretti & Shima, 2011.—Afrotropical: D.R. Congo.
Dolichocolon bequaerti Cerretti & Shima, 2011: 556. Holotype male (MRAC). Type locality: D.R. Congo, Katanga, Kunda (7°15′S 28°27′E).
caudatum Cerretti & Shima, 2011.—Afrotropical: Senegal.
Dolichocolon caudatum Cerretti & Shima, 2011: 561. Holotype male (SMNS). Type locality: Senegal, Simenti (13°02′N 13°18'W), Maribor.
crosskeyi Cerretti & Shima, 2011.—Afrotropical: Angola, Zimbabwe.
Dolichocolon crosskeyi Cerretti & Shima, 2011: 565. Holotype male (BMNH). Type locality: Zimbabwe, Chikurubi (17°47′S 31°12′E).
elegans Cerretti & Shima, 2011.—Afrotropical: D.R. Congo.
Dolichocolon elegans Cerretti & Shima, 2011: 553. Holotype male (MRAC). Type locality: D.R. Congo, Katanga, Lubumbashi.
meii Cerretti & Shima, 2011.—Afrotropical: Ethiopia.
Dolichocolon meii Cerretti & Shima, 2011: 554. Holotype male (MZUR). Type locality: Ethiopia, El Banno, 1250m (4°51′0.05″N 37°23′59.96″E).
mesnili Cerretti & Shima, 2011.—Afrotropical: D.R. Congo.
Dolichocolon mesnili Cerretti & Shima, 2011: 560. Holotype male (CNC). Type locality: D.R. Congo, Katanga, Lubumbashi.
paradoxum Brauer & Bergenstamm, 1889.—Afrotropical: D.R. Congo, Mozambique. Palaearctic: Europe (W. Eur., SW. Eur., SC. Eur., SE. Eur.), M. East (all), Transcaucasia.
Dolichocolon paradoxum Brauer & Bergenstamm, 1889: 100, 165 [also 1890: 32, 97]. Holotype male [not lectotype male as inferred by O’Hara et al. 2009: 106, see Cerretti and Shima 2011: 555] (NHMW). Type locality: Croatia, Dalmacija [as “Dalmatien”].
Note: Cerretti and Shima (2011: 555–556) redescribed Dolichocolon paradoxum Brauer & Bergenstamm, 1889 and reevaluated its distribution. These authors noted that D. paradoxum had been misidentified from South Africa in Crosskey (1980b: 877) and its presence in eastern Asia as recorded in O’Hara et al. (2009: 106) and elsewhere is suspect (Cerretti and Shima 2011: 556).
paravicinum Cerretti & Shima, 2011.—Afrotropical: Nigeria, South Africa, Yemen.
Dolichocolon paravicinum Cerretti & Shima, 2011: 571. Holotype male (RMNH). Type locality: Yemen, 12km northwest of Manākhah (15°04′19″N 43°44′27″E according to Zeegers 2009: 371).
Note: The holotype of Dolichocolon paravicinum Cerretti & Shima, 2011 is one of the specimens from Yemen that Zeegers (2007: 384) examined and cited as “Dolichocolon sp.”. Specimens of D. paravicinum from Uganda were misidentified as Dolichocolon vicinum Mesnil, 1968 in Crosskey (1980b: 877). Dolichocolon vicinum is currently regarded as a strictly Oriental species (Cerretti and Shima 2011: 569–571).
rude Cerretti & Shima, 2011.—Afrotropical: Cameroon, Côte d’Ivoire, D.R. Congo, South Africa.
Dolichocolon rude Cerretti & Shima, 2011: 558. Holotype male (NMDA). Type locality: Cameroon, Kassei (10°31′N 14°46′E).
Genus ERYTHROCERA Robineau-Desvoidy, 1849
ERYTHROCERA Robineau-Desvoidy, 1848: 186. Nomen nudum (no description or included species).
ERYTHROCERA Robineau-Desvoidy, 1849b: 436. Type species: Phryno nigripes Robineau-Desvoidy, 1830, by subsequent designation of Robineau-Desvoidy (1863a: 600, as “Erythrocera nigripes, R.-D.”) [Palaearctic].
doris (Curran, 1927).—Afrotropical: D.R. Congo.
Sturmia doris Curran, 1927c: 18 (as “Sturmia (?) doris”). Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
Sturmia dorina Curran, 1927f: 126 (unnecessary nomen novum for Sturmia doris Curran, 1927).
Note: Sturmia dorina Curran, 1927 was proposed for “Sturmia ? doris Curran (not Schiner)”. However, this was based on the misidentification of Tachina doris Meigen, 1824 by previous authors. Herting (1972: 5) established the true T. doris Meigen as a junior synonym of Lydella stabulans (Meigen, 1824) (see also Herting and Dely-Draskovits 1993: 205 [doris Meigen], 248 [doris of authors, not Meigen]).
picta (Villeneuve, 1936).—Afrotropical: Nigeria.
Pexomyia (Erythrocera) picta Villeneuve, 1936a: 7. Holotype male (CNC). Type locality: Nigeria, Oshogbo.
porcula Mesnil, 1952.—Afrotropical: Nigeria, Sierra Leone.
Erythrocera porcula Mesnil, 1952c: 252. Holotype female (not located). Type locality: northern Nigeria.
Genus GONIA Meigen, 1803
SALMACIA Meigen, 1800: 38. Name suppressed by ICZN (1963: 339).
GONIA Meigen, 1803: 280. Type species: Gonia bimaculata Wiedemann, 1819, by subsequent designation of Sabrosky and Arnaud (1965: 1075).
bimaculata Wiedemann, 1819.—Afrotropical: “widespread mainland Afrotrop. Reg. (excl. W. Afr.)” (Crosskey 1980b: 875), including Malawi, South Africa, Uganda, Yemen. Palaearctic: C. Asia, Europe (E. Eur., SW. Eur., SC. Eur., SE. Europe, Turkey), M. East (all), N. Africa (Canary Is., Madeira), Pal. China, Transcaucasia. Oriental: Orien. China.
Gonia bimaculata Wiedemann, 1819: 25. Type(s), female (1 female in NHMW, 1 syntype in ZMUC [Zimsen 1954: 23]). Type locality: South Africa, Western Cape, Cape of Good Hope [as “Prom. bon. sp.” = “Promontorium Bonae Spei”].
Note: Gonia bimaculata Wiedemann, 1819 was described from an unspecified number of females, but certainly more than one because Wiedemann (1930: 344) later wrote “In Westermann’s und meiner Sammlung”. A female in NHMW (examined by JEOH) is recognizable as a syntype by its “bimaculata Coll. Wiedem.” label and a second label giving the species name, locality (“C. b. sp.”), and collector (“Westermann”). A third label, blue with only “Typus” handwritten on it, appears to have been written by Villeneuve.
rubriventris Macquart, 1851.—Afrotropical: South Africa.
Gonia rubriventris Macquart, 1851b: 150 [also 1851b: 177]. Lectotype female (BMNH), by fixation of Crosskey (1971: 270) (examination of “Holotype ♀” from Cape of Good Hope in BMNH is regarded as a lectotype fixation). Type locality: South Africa, Western Cape, Cape of Good Hope [as “Cap de Bonne-Espérance”].
Note: Crosskey (1984: 285) discussed the differences between Gonia rubriventris Macquart, 1851 and Gonia bimaculata Wiedemann, 1819 and suggested they are “possibly synonymous”.
Genus GONIOPHTHALMUS Villeneuve, 1910
GONIOPHTHALMUS Villeneuve in Becker, 1910b: 145 [also 1910b: 15]. Type species: Goniophthalmus simonyi Villeneuve, 1910, by monotypy.
halli Mesnil, 1956.—Afrotropical: Botswana, Cape Verde, Kenya, Namibia, Sudan, Tanzania, U.A. Emirates, Yemen, Zimbabwe. Palaearctic: M. East (all). Oriental: India.
Goniophthalmus halli Mesnil, 1956c: 548. Holotype male (published as BMNH but probably not deposited there according to Crosskey 1976: 244). Type locality: Zimbabwe, Mazoe.
simonyi Villeneuve. 1910.—Afrotropical: Yemen.
Goniophthalmus simonyi Villeneuve in Becker, 19l0b: 145 [also 1910b: 15]. Lectotype male (NHMW), by fixation of Townsend (1941: 34) (mention of “Ht male” from “Ras Shoab, Sokotra” in NHMW is regarded as a lectotype fixation for the single male in the type series). Type locality: Yemen, Suquţrá [as “Sokotra”] (“Ras Shoab” according to Townsend 1941: 34 and label data of lectotype [examined by JEOH]).
Undescribed sp.: Kenya (MZUR, examined by PC).
Genus HYSTRICEPHALA Macquart, 1846
HYSTRICEPHALA Macquart, 1846: 282 [also 1846: 154]. Type species: Hystricephala nigra Macquart, 1846, by monotypy.
nigra Macquart, 1846.—Afrotropical: South Africa.
Hystricephala nigra Macquart, 1846: 283 [also 1846: 155]. Holotype male (“presumed lost”, Crosskey 1971: 272). Type locality: South Africa, “Cafrerie” (also as “Kaffraria”; probably referring to an area now in the southeastern part of Eastern Cape).
Genus IGNEOMYIA Mesnil, 1950
IGNEOMYIA Mesnil, 1949b: 103 (as subgenus of Congochrysosoma Townsend, 1916). Nomen nudum (proposed after 1930 without designation of type species; no included species) (see Evenhuis and O’Hara 2008: 66).
IGNEOMYIA Mesnil, 1950c: 105, 108 (as subgenus of Congochrysosoma Townsend, 1916). Type species: Pexopsis (Ugimeigenia) ignea Mesnil, 1944, by monotypy (see Evenhuis and O’Hara 2008: 66).
ferruginea Mesnil, 1970.—Afrotropical: Madagascar.
Igneomyia ferruginea Mesnil, 1970b: 107. Holotype male (MNHN). Type locality: Madagascar, Antsiranana, Montagne d’Ambre [Parc National, ca. 12°36′S 49°8′E].
ignea (Mesnil, 1944).—Afrotropical: Madagascar.
Pexopsis (Ugimeigenia) ignea Mesnil, 1944: 10. Holotype male (MNHN). Type locality: Madagascar, Toliara, Bekily.
Genus KUWANIMYIA Townsend, 1916
KUWANIMYIA Townsend, 1916d: 319. Type species: Kuwanimyia conspersa Townsend, 1916, by original designation [Palaearctic].
Note: Kuwanimyia Townsend, 1916 was revised by Cerretti (2009b).
afra Cerretti, 2009.—Afrotropical: Namibia.
Kuwanimyia afra Cerretti, 2009b: 56. Holotype male (BMNH). Type locality: Namibia, 23 miles southwest of Grootfontein, Rietfontein.
atra Cerretti, 2009.—Afrotropical: Namibia, Nigeria.
Kuwanimyia atra Cerretti, 2009b: 57. Holotype male (BMNH). Type locality: Nigeria, Samaru.
capensis Cerretti, 2009.—Afrotropical: South Africa.
Kuwanimyia capensis Cerretti, 2009b: 58. Holotype female (NMDA). Type locality: South Africa, Eastern Cape, Fort Beaufort.
Genus LYDELLINA Villeneuve, 1916
LYDELLINA Villeneuve, 1916c: 490. Type species: hereby fixed under Article 70.3.2 of the Code (ICZN 1999) as Lydellina villeneuvei Townsend, 1933, misidentified as Masicera caffra Macquart, 1846 in the fixation by monotypy of Villeneuve (1916c).
anorbitalis Mesnil, 1970.—Afrotropical: Benin, Tanzania, Uganda.
Lydellina anorbitalis Mesnil, 1970b: 99. Holotype male (CNC). Type locality: Benin [as “Dahomey”], Agouagon [ca. 7°59′N 2°18′E].
distincta Mesnil, 1970.—Afrotropical: Madagascar.
Lydellina distincta Mesnil, 1970b: 100. Holotype male (MNHN). Type locality: Madagascar, Toliara, Bekily.
frontalis Mesnil, 1970.—Afrotropical: Ghana.
Lydellina frontalis Mesnil, 1970b: 100. Holotype male (CNC). Type locality: Ghana, Aburi.
umbripennis Mesnil, 1970.—Afrotropical: D.R. Congo.
Lydellina umbripennis Mesnil, 1970b: 100. Holotype male (CNC). Type locality: D.R. Congo, Équateur, Eala.
villeneuvei Townsend, 1933.—Afrotropical: D.R. Congo, Malawi, South Africa.
Lydellina villeneuvei Townsend, 1933: 469 (named for caffra of Villeneuve, 1916c, etc., not Macquart, 1846). Holotype female (SAMC). Type locality: South Africa, KwaZulu-Natal, Durban.
Masicera caffra of authors (e.g., Villeneuve 1916c: 490, Curran 1928b: 397, Verbeke 1962b: 50, all three as “Lydellina caffra”), not Macquart, 1846. Misidentification (Crosskey 1980b: 879).
Genus MINTHOSOMA Zeegers, 2007
MINTHOSOMA Zeegers, 2007: 389. Type species: Minthosoma janus Zeegers, 2007, by original designation.
Note: We have followed Zeegers (2007: 389) in tentatively placing this genus in the Goniini (“genus is likely to be close to Baumhaueria [Meigen]”).
janus Zeegers, 2007.—Afrotropical: Yemen.
Minthosoma janus Zeegers, 2007: 390. Holotype female (RMNH). Type locality: Yemen, Seyun (15°56′36″N 48°47′36″E).
Genus MYXOGAEDIA Mesnil, 1956
PRETORIANA Curran, 1938: 7 (junior homonym of Pretoriana Uvarov, 1922). Type species: Pretoriana setosa Curran, 1938, by original designation.
MYXOGAEDIA Mesnil, 1956a: 497. Type species: Myxarchiclops maculosus Villeneuve, 1916, by original designation.
GAUTENGICESA Koçak & Kemal, 2010: 157 (nomen novum for Pretoriana Curran, 1938).
Note: Myxogaedia Mesnil, 1956 was recognized as the valid name for this genus by O’Hara (2011: 60–61) after Gautengicesa Koçak & Kemal, 2010 was proposed as a replacement name for Pretoriana Curran, 1938.
maculosa (Villeneuve, 1916).—Afrotropical: South Africa.
Myxarchiclops maculosus Villeneuve, 1916c: 496 (as “Myxarchiclops (?) maculosus”). Holotype female (CNC). Type locality: South Africa, Northern Cape, Springbok [as “Springbokfontein”].
setosa (Curran, 1938).—Afrotropical: South Africa.
Pretoriana setosa Curran, 1938: 7. Holotype male (SANC). Type locality: South Africa, Gauteng, Pretoria.
Undetermined sp. (nr. M. maculosa (Villeneuve)): Namibia (MZUR, examined by PC).
Genus MYXOPHRYXE Cerretti & O’Hara, gen. n.
MYXOPHRYXE Cerretti & O’Hara, gen. n. Type species: Phorocera longirostris Villeneuve, 1938, by present designation.
Note: This new genus and the three new species below are described in the New Taxa of Afrotropical Tachinidae section.
longirostris (Villeneuve, 1938).—Afrotropical: South Africa. Comb. n.
Phorocera longirostris Villeneuve, 1938c: 2. Holotype male (not located; male specimen in CNC labelled by Mesnil as “TYPE” and cited as such by Cooper and O’Hara 1996: 62 is not from the type locality and is not the holotype). Type locality: South Africa, “Colonie du Cap” (former Cape Province, corresponding to the present-day Western Cape, Eastern Cape, Northern Cape, and North West [in part] provinces).
Phorocera majestica Curran, 1940: 10. Holotype male (SANC). Type locality: South Africa, KwaZulu-Natal, New Hanover. Syn. n.
Note: Phorocera longirostris Villeneuve, 1938 and Phorocera majestica Curran, 1940 were treated as species of Pretoriana Curran, 1938 (the valid name of which is now Myxogaedia Mesnil, 1956) by Crosskey (1980b: 879). These nominal species are moved here to Myxophryxe gen. n., with P. longirostris as the valid name and P. majestica in synonymy. This species is redescribed in the New Taxa of Afrotropical Tachinidae section.
murina Cerretti & O’Hara, sp. n.—Afrotropical: South Africa.
Myxophryxe murina Cerretti & O’Hara, sp. n. Holotype male (NMB). Type locality: South Africa, Western Cape, De Vasselot Natural Reserve (33°58.194′S 23°32.193′E).
regalis Cerretti & O’Hara, sp. n.—Afrotropical: South Africa.
Myxophryxe regalis Cerretti & O’Hara, sp. n. Holotype male (NMB). Type locality: South Africa, KwaZulu-Natal, Royal Natal National Park, Thendele, 1600m (28°42.378′S 28°56.083′E).
satanas Cerretti & O’Hara, sp. n.—Afrotropical: South Africa.
Myxophryxe satanas Cerretti & O’Hara, sp. n. Holotype male (MZUR). Type locality: South Africa, Western Cape, Gamkaskloof (Die Hel), 336m (33°21′49.60″S 21°37′40.97″E).
Genus NEALSOMYIA Mesnil, 1939
NEALSOMYIA Mesnil, 1939d: 31. Type species: Exorista (Alsomyia) triseriella Villeneuve, 1929, by original designation [Palaearctic].
Note: A world revision of Nealsomyia Mesnil, 1939 was published by Cerretti (2005).
chloronitens (Mesnil, 1977).—Afrotropical: Madagascar. Comb. n.
Alsomyia chloronitens Mesnil, 1977b: 187. Holotype male (MNHN). Type locality: Madagascar, Ambohitantely [Réserve Spéciale, ca. 18°10′S 47°17′E].
Note: Alsomyia Brauer & Bergenstamm, 1891 was recognized from the Afrotropical Region by Mesnil (1977b) based on his new species A. chloronitens Mesnil, 1977. This species is moved here to Nealsomyia Mesnil, 1939.
clausa (Curran, 1940).—Afrotropical: Zimbabwe. Comb. n.
Phorocera clausa Curran, 1940: 9. Holotype male (AMNH). Type locality: Zimbabwe, Kadoma [as “Gatooma”].
Note: Phorocera clausa Curran, 1940 was treated as an unplaced species of “Goniinae” [= Exoristinae] by Crosskey (1980b: 881) but is moved here to Nealsomyia Mesnil, 1939 based on examination of the holotype.
lindneri Mesnil, 1959.—Afrotropical: Tanzania.
Nealsomyia lindneri Mesnil, 1959: 12. Holotype male (SMNS). Type locality: Tanzania, Lake Victoria, Mugango.
merzi Cerretti, 2005.—Afrotropical: Namibia.
Nealsomyia merzi Cerretti, 2005: 129. Holotype male (MHNG). Type locality: Namibia, Mount Erongo.
Undescribed sp.: South Africa (NMB, examined by PC).
Genus PALES Robineau-Desvoidy, 1830
PALES Robineau-Desvoidy, 1830: 154 (not a junior homonym of Pales Meigen, 1800 [Tipulidae] because the work in which that name appeared was suppressed by ICZN 1963: 339). Type species: Pales florea Robineau-Desvoidy, 1830 (= Tachina pavida Meigen, 1824), by subsequent designation of Coquillett (1910: 582) [Palaearctic].
CTENOPHOROCERA Brauer & Bergenstamm, 1891: 342 [also 1891: 38]. Type species: Ctenophorocera blepharipus Brauer & Bergenstamm, 1891, by subsequent designation of Sharp (1893: 299).
NEOPALES Coquillett, 1910: 575 (nomen novum for Pales Robineau-Desvoidy, 1830; proposed prior to the suppression of Pales Meigen, 1800 by ICZN 1963: 339).
MICROPALES Villeneuve, 1927: 121. Type species: Micropales seminitida Villeneuve, 1927, by monotypy.
aethiopica (Mesnil, 1950).—Afrotropical: D.R. Congo, South Africa, Sudan, Tanzania.
Ctenophorocera (Ctenophorocera) aethiopica Mesnil, 1950c: 124. Holotype male (CNC). Type locality: northwestern Tanzania, forest edge, 1800–2000m.
blepharipa (Brauer & Bergenstamm, 1891).—Afrotropical: D.R. Congo, South Africa, Uganda.
Ctenophorocera blepharipus Brauer & Bergenstamm, 1891: 342 [also 1891: 38]. Type(s), male (NHMW, not located by JEOH). Type locality: South Africa, Western Cape, Cape of Good Hope [as “?(Cap oder Brasilien. Coll. Winth.)”].
Note: Aldrich (1927d: 24) examined a male in NHMW bearing “a large blue name label in Doctor Villeneuve’s writing” that he treated as the “type” of Ctenophorocera blepharipus Brauer & Bergenstamm, 1891. The specimen was also labelled “caffra. Coll. Winthem”, with “caffra” meaning “Caffraria”, a former region in Eastern Cape (also known as “Kaffraria”). This is a different locality from the two possible localities given in the original description (Cape of Good Hope or Brazil). Given the uncertainty about whether this specimen is a name-bearing type of C. blepharipus, Aldrich’s (1927d: 24) examination of the “type” is not accepted as a lectotype fixation. JEOH did not find a specimen in NHMW that matches the expected type data for C. blepharipus.
coerulea (Jaennicke, 1867).—Afrotropical: “n.-e. Afr. to sthn Afr.” (Crosskey 1980b: 870), including Ethiopia, South Africa, Zimbabwe.
Phorocera coerulea Jaennicke, 1867: 382 [also 1868: 74]. Type(s), male (SMF). Type locality: Ethiopia, “Simen” (probably the Simien Mountains area).
caerulea. Incorrect subsequent spelling of coerulea Jaennicke, 1867 (Aldrich 1927: 23).
Note: Crosskey (1980b: 870) noted the possible presence of Phorocera coerulea Jaennicke, 1867 in the Oriental Region, probably based on Mesnil’s (1950c: 126) mention of “?Indien”. The presence of this species in the Oriental Region needs confirmation.
coeruleonigra (Mesnil, 1950).—Afrotropical: Zimbabwe.
Ctenophorocera (Ctenophorocera) coerulea coeruleonigra Mesnil, 1950c: 126 (as “coeruleo-nigra”). Holotype male (CNC). Type locality: Zimbabwe, Mutare [as “Umtali”] District, Vumba Mountains.
contristans Villeneuve, 1938.—Afrotropical: South Africa.
Pales contristans Villeneuve, 1938c: 1. Type(s), unspecified sex (1 male in CNC). Type locality: South Africa, “Colonie du Cap” ([former Cape Province], Algoa Bay, according to label data of CNC syntype, Cooper and O’Hara 1996: 57).
corrupta (Curran, 1927).—Afrotropical: Uganda.
Zenillia corrupta Curran, 1927d: 331. Holotype male (BMNH). Type locality: Uganda, Jeza [ca. 0°22′N 32°17′E].
Note: Curran (1927d: 331) reported that the holotype of his new species Zenillia corrupta was reared from a syrphid (Diptera, Syrphidae). This is a dubious record, but if true would be the only known case of parasitism of a syrphid (larva?) by a tachinid.
cuthbertsoni (Curran, 1940).—Afrotropical: Zimbabwe.
Phorocera cuthbertsoni Curran, 1940: 5. Holotype male (AMNH). Type locality: Zimbabwe, Nyanga [as “Inyanga”].
cuthbersoni. Incorrect subsequent spelling of cuthbertsoni Curran, 1940 (original usage not found but spelling listed by Crosskey 1980b: 870).
divergens (Curran, 1928).—Afrotropical: Uganda.
Phorocera divergens Curran, 1928a: 237. Holotype male (BMNH). Type locality: Uganda, Kampala.
experta (Brauer & Bergenstamm, 1891).—Afrotropical: South Africa.
Ctenophorocera experta Brauer & Bergenstamm, 1891: 342 [also 1891: 38] (as “experta Wd.”). Lectotype male (NHMW, not located by JEOH), by fixation of Townsend (1941: 98) (mention of “Ht male” from Cape of Good Hope in NHMW is regarded as a lectotype fixation). Type locality: South Africa, Western Cape, Cape of Good Hope [as “Cap b. sp.” = “Cap Bonae Spei”].
gnu (Curran, 1940).—Afrotropical: Liberia, Nigeria, Rwanda.
Phorocera gnu Curran, 1940: 11. Holotype male (AMNH). Type locality: Liberia, Ganta.
macrocephala (Mesnil, 1950).—Afrotropical: Kenya (new record, MZUR [PC]), Malawi, South Africa.
Ctenophorocera (Ctenophorocera) macrocephala Mesnil, 1950c: 123. Holotype male (CNC). Type locality: Malawi, Nyika Plateau.
maculisquama (Mesnil, 1950).—Afrotropical: Zimbabwe.
Ctenophorocera (Ctenophorocera) coerulea maculisquama Mesnil, 1950c: 126. Holotype male (CNC). Type locality: Zimbabwe, Harare [as “Salisbury”].
Note: Crosskey (1980b: 870) noted the possible presence of Ctenophorocera coerulea maculisquama Mesnil, 1950 in the Oriental Region, probably based on Mesnil’s (1950c: 126) statement: “?. Ein defektes ♂ aus Indien”. The presence of this species in the Oriental Region needs confirmation.
metro (Curran, 1940).—Afrotropical: Zambia, Zimbabwe.
Phorocera metro Curran, 1940: 12. Holotype male (AMNH). Type locality: border between Zambia and Zimbabwe, Victoria Falls.
nigronitens Villeneuve, 1938.—Afrotropical: D.R. Congo, South Africa.
Pales nigronitens Villeneuve, 1938c: 1 (as “nigro-nitens”). Syntypes, males and females (?IRSNB). Type locality: D.R. Congo.
Phorocera ethelia Curran, 1940: 9. Holotype male (AMNH). Type locality: South Africa, KwaZulu-Natal, Durban.
nyasa (Curran, 1940).—Afrotropical: Malawi, South Africa.
Phorocera nyasa Curran, 1940: 13. Holotype male (BMNH). Type locality: Malawi, Nsanje [as “Port Herald”].
pauciseta (Mesnil, 1950).—Afrotropical: D.R. Congo.
Ctenophorocera (Ctenophorocera) pauciseta Mesnil, 1950c: 125. Holotype male (CNC). Type locality: D.R. Congo, Équateur, Eala (see O’Hara and Cooper 1996: 27 for label data).
Note: O’Hara (1996: 152) commented on the holotype of Ctenophorocera pauciseta Mesnil, 1950: “The type locality is stated as Kisantu but the specimen labelled as the holotype is from Eala. The length of the Eala specimen corresponds with the length given in the description, whereas the two CNC specimens from Kisantu are larger.”
rubrica Villeneuve, 1932.—Afrotropical: Kenya, Tanzania.
Pales rubrica Villeneuve, 1932: 285. Holotype male (BMNH). Type locality: Kenya, Aberdare Mountains, 7300ft.
rubriventris Bezzi, 1908.—Afrotropical: South Africa.
Pales rubriventris Bezzi, 1908a: 185. Holotype female (not located). Type locality: South Africa, Northern Cape, Steinkopf.
ruficauda (Curran, 1927).—Afrotropical: D.R. Congo.
Phorocera ruficauda Curran, 1927c: 9. Holotype female (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
rufolateralis (Curran, 1940).—Afrotropical: Kenya, Malawi, South Africa.
Phorocera rufolateralis Curran, 1940: 11. Holotype male (BMNH). Type locality: Malawi, Zomba.
sarcophagaeformis (Jaennicke, 1867).—Afrotropical: Ethiopia, Kenya, Malawi, South Africa, Tanzania, Uganda.
Phorocera sarcophagaeformis Jaennicke, 1867: 381 [also 1868: 73]. Type(s), male (SMF). Type locality: Ethiopia, “Simen” (probably the Simien Mountains area).
seminitida (Villeneuve, 1927).—Afrotropical: D.R. Congo, Malawi, Nigeria, Zimbabwe.
Micropales seminitida Villeneuve, 1927: 121. Lectotype male (BMNH), by fixation of Townsend (1941: 108) (mention of “Ht male” from Ibadan in BMNH is regarded as a lectotype fixation). Type locality: Nigeria, Ibadan.
senex (Curran, 1927).—Afrotropical: D.R. Congo, Nigeria.
Phorocera senex Curran, 1927c: 10. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
setigena (Curran, 1940).—Afrotropical: South Africa, Zimbabwe.
Phorocera setigena Curran, 1940: 11. Holotype male (BMNH). Type locality: South Africa, KwaZulu-Natal, Stella Bush [near Durban].
Note: Curran (1940: 11) cited the type locality of Phorocera setigena as “Marley, Stella Bush”, but Marley was the collector.
somomyina (Karsch, 1886).—Afrotropical: Angola.
Phorocera somomyina Karsch, 1886b: 340. Holotype, unspecified sex (ZMHB, not located by JEOH). Type locality: Angola, Pungo Andongo.
splendens Mesnil, 1970.—Afrotropical: Madagascar.
Pales splendens Mesnil, 1970b: 89. Holotype male (MNHN). Type locality: Madagascar, Toamasina, Moramanga.
tessellans (Mesnil, 1950).—Afrotropical: South Africa.
Ctenophorocera (Ctenophorocera) tessellans Mesnil, 1950c: 123. Holotype male (CNC). Type locality: South Africa, KwaZulu-Natal.
tetra (Curran, 1940).—Afrotropical: South Africa.
Phorocera tetra Curran, 1940: 12. Holotype female (SANC). Type locality: South Africa, Mpumalanga, Barberton.
Genus PERLUCIDINA Mesnil, 1952
PERLUCIDINA Mesnil, 1949b: 104 (as subgenus of Tamaromyia Mesnil, 1949). Nomen nudum (proposed after 1930 without designation of type species; no included species) (see Evenhuis and O’Hara 2008: 67).
PERLUCIDINA Mesnil, 1952c: 223 (as subgenus of Hygia Mesnil, 1952 [not Hygia Uhler, 1861]). Type species: Exorista perlucida Karsch, 1886, by monotypy (see Evenhuis and O’Hara 2008: 67). Status revived.
Note: Crosskey (1980b: 869) synonymized Perlucidina Mesnil, 1952 with Calozenillia Townsend, 1927. We do not agree with this synonymy and here reinstate Perlucidina as a genus. The characters that distinguish Perlucidina will be given in the Tachinidae chapter of the Manual of Afrotropical Diptera (in prep.).
africana (Jaennicke, 1867).—Afrotropical: Ethiopia. Comb. n.
Exorista africana Jaennicke, 1867: 384 [also 1868: 76]. Type(s), female (SMF). Type locality: Ethiopia, “Simen” (probably the Simien Mountains area).
Note: Exorista africana Jaennicke, 1867 was treated as a species of Calozenillia Townsend, 1927 by Crosskey (1980b: 869, 1984: 281) but is moved here to the newly revived genus Perlucidina Mesnil, 1952.
perlucida (Karsch, 1886).—Afrotropical: Angola, D.R. Congo, Malawi, South Africa, Sudan, Uganda, Zambia. Comb. revived.
Tachina dubia Walker, 1853: 291 (junior primary homonym of Tachina dubia Fallén, 1810). Type(s), female (BMNH). Type locality: South Africa, Western Cape, Cape of Good Hope [as “Cape”].
Exorista perlucida Karsch, 1886b: 339. Holotype, unspecified sex [male, examined by JEOH] (ZMHB). Type locality: Angola, Pungo Andongo.
Note: Exorista perlucida Karsch, 1886b was treated as a species of Calozenillia Townsend, 1927 by Crosskey (1980b: 869, 1984: 281) but is moved here to the newly revived genus Perlucidina Mesnil, 1952.
Genus PEXOPSIS Brauer & Bergenstamm, 1889
PEXOPSIS Brauer & Bergenstamm, 1889: 88 [also 1890: 20]. Type species: Eurigaster tibialis Robineau-Desvoidy, 1849 (as “tibialis Mg.”) (= Tachina aprica Meigen, 1824), by monotypy [Palaearctic].
chapini (Curran, 1927).—Afrotropical: D.R. Congo, Kenya, Uganda.
Sturmia chapini Curran, 1927a: 11. Holotype male (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
femoralis Bezzi, 1911.—Afrotropical: Malawi, Mozambique.
Pexopsis femoralis Bezzi, 1911: 59. Holotype female (USNM). Type locality: Mozambique, Maputo Province, “Umbelusi”.
garambana Verbeke, 1962.—Afrotropical: D.R. Congo.
Pexopsis garambana Verbeke, 1962b: 51. Holotype female (IRSNB). Type locality: D.R. Congo, Orientale, Parc National de la Garamba [as “P.N.G.”].
lindneri Mesnil, 1959.—Afrotropical: D.R. Congo, Tanzania.
Pexopsis lindneri Mesnil, 1959: 10. Holotype male (SMNS). Type locality: Tanzania, Pare Mountains, 1800m.
pyrrhaspis Villeneuve, 1916.—Afrotropical: “widespread W. Afr., E. Afr. & sthn Afr.” (Crosskey 1980b: 873), including Kenya, Malawi, South Africa.
Pexopsis pyrrhaspis Villeneuve, 1916c: 492. Syntypes, 2 females (BMNH, SAMC). Type localities: Malawi and South Africa, “Cape Colony” (former Cape Province, corresponding to the present-day Western Cape, Eastern Cape, Northern Cape, and North West [in part] provinces).
yemenensis Zeegers, 2007.—Afrotropical: Yemen.
Pexopsis yemenensis Zeegers, 2007: 393. Holotype male (RMNH). Type locality: Yemen, Wādī Lahīmah [as “Al Lahima”] (15°24′N 43°32′E).
Genus PHYTOMYPTERINA van Emden, 1960
PHYTOMYPTERINA van Emden, 1960: 356. Type species: Phytomypterina burtti van Emden, 1960 (= Phytomyptera rufescens Villeneuve, 1936), by original designation.
rufescens (Villeneuve, 1936).—Afrotropical: Mozambique (new record, MZUR [PC]), South Africa, Tanzania.
Phytomyptera rufescens Villeneuve, 1936a: 3. Holotype female (CNC). Type locality: South Africa (KwaZulu-Natal, Mfongosi [as “M fongosi Zulu L.”] according to label data, Cooper and O’Hara 1996: 63).
Phytomypterina burtti van Emden, 1960: 357. Holotype male (BMNH). Type locality: Tanzania, Singida.
Genus PIMELIMYIA Mesnil, 1949
PIMELIMYIA Mesnil, 1949b: 104. Type species: Sturmia russata Villeneuve, 1934 (as “Pimelimyia russata Vill.”, p. 103), by monotypy (see Evenhuis and O’Hara 2008: 67).
grossa Mesnil, 1959.—Afrotropical: Tanzania, Zimbabwe.
Pimelimyia grossa Mesnil, 1959: 10. Holotype female (SMNS). Type locality: Tanzania, Pare Mountains, Usangi.
insularis (Villeneuve, 1915).—Afrotropical: Madagascar.
Sturmia insularis Villeneuve, 1915b: 193. Syntypes, 1 male and 1 female (male in NHMW, female not located). Type localities: Madagascar, female syntype from Antananarivo, Antananarivo [as “Tananarive”], male syntype (examined by JEOH) from “Mgdk.” [= Madagascar] without further locality data.
Note: Sturmia insularis Villeneuve, 1915 is probably misplaced in Pimelimyia Mesnil, 1949; it lacks a sexual patch on the underside of abdominal tergite 4 but is otherwise similar to species of Blepharipa Rondani, 1856 (a genus not recorded from the Afrotropical Region).
natalensis (Curran, 1927).—Afrotropical: South Africa.
Sturmia natalensis Curran, 1927f: 121. Holotype female (BMNH). Type locality: South Africa, KwaZulu-Natal, Weenen [ca. 28°51′S 30°4′E].
rufina (Curran, 1927).—Afrotropical: South Africa.
Sturmia rufina Curran, 1927f: 125. Holotype female (SANC). Type locality: South Africa, Gauteng, Pretoria.
rufula (Villeneuve, 1943).—Afrotropical: “Afrique orientale” [East Africa], South Africa.
Sturmia rufula Villeneuve, 1943a: 38. Syntypes, 2 females (not located). Type locality: “Afrique orientale” [East Africa].
russata (Villeneuve, 1943).—Afrotropical: “l’Afrique orientale” [East Africa], South Africa.
Sturmia russata Villeneuve, 1943a: 37. Syntypes, 2 males (not located). Type localities: South Africa and “l’Afrique orientale” [East Africa].
semitestacea (Villeneuve, 1916).—Afrotropical: Malawi, South Africa, Tanzania, Zimbabwe.
Sturmia (Blepharipoda) semitestacea Villeneuve, 1916c: 477. Syntypes, 7 males and females (BMNH, SAMC [1 male examined by JEOH]). Type localities: Malawi and South Africa [latter inferred from deposition of specimens in “S. Afric. Museum”].
Genus PROSOPODOPSIS Townsend, 1926
PROSOPODOPSIS Townsend, 1926b: 542. Type species: Tachina fasciata Wiedemann, 1830 (junior primary homonym of Tachina fasciata Fallén, 1820; = Prosopaea appendiculata de Meijere, 1910), by original designation [Oriental].
Note: Three undescribed species from Namibia, Nigeria and Uganda assigned to Prosopodopsis Townsend, 1926 by Crosskey (1980b: 880, 1984: 295) were placed elsewhere by Cerretti (2009b: 52).
pulchricornis (Villeneuve, 1938).—Afrotropical: Mozambique, ?South Africa.
Histochaeta pulchricornis Villeneuve, 1938a: 3. Holotype male (CNC). Type locality: Africa (“Erosba Pan” according to label data [an unknown location], collected by the “S.W. Africa Mus. Exped.”); considered “sud-africaine très probablement” by Villeneuve (1938a: 3).
Note: Histochaeta pulchricornis Villeneuve, 1938 was treated as a species of Chlorolydella Townsend, 1933 by Crosskey (1980b: 877) but was moved to Prosopodopsis Townsend, 1926 by Crosskey (1984: 201, 295). This placement was upheld by Cerretti (2009b: 52).
Undescribed sp.: Nigeria (CNC, examined by PC).
Undetermined spp.: U.A. Emirates (Zeegers 2010), Yemen (Zeegers 2007: 394).
Genus PSEUDALSOMYIA Mesnil, 1968
PSEUDALSOMYIA Mesnil, 1968c: 178. Type species: Pseudalsomyia piligena Mesnil, 1968, by original designation [Oriental].
audisioi Cerretti, 2012.—Afrotropical: Kenya.
Pseudalsomyia audisioi Cerretti, 2012: 329. Holotype male (MZUR). Type locality: Kenya, Western, Kakamega Forest, 1600m (0°13′37.2″N 34°52′49.8″E).
Genus PSEUDOGONIA Brauer & Bergenstamm, 1889
PSEUDOGONIA Brauer & Bergenstamm, 1889: 100 [also 1890: 32]. Type species: Gonia cinerascens Rondani, 1859 (= Tachina rufifrons Wiedemann, 1830), by monotypy [Palaearctic].
GAEDIOGONIA Townsend, 1927c: 71. Type species: Gaediogonia jacobsoni Townsend, 1927 (= Tachina rufifrons Wiedemann, 1830), by original designation [Oriental].
fasciata (Wiedemann, 1819).—Afrotropical: South Africa, Zimbabwe. Palaearctic: Europe (SW. Eur.), N. Africa (Canary Is.).
Gonia fasciata Wiedemann, 1819: 25. Syntypes, female (2 syntypes in ZMUC, Zimsen 1954: 23). Type locality: South Africa, Western Cape, Cape of Good Hope [as “Prom. bon. sp.” = “Promontorium Bonae Spei”].
Rhedia capensis Robineau-Desvoidy, 1830: 77. Type(s), unspecified sex (MNHN or lost). Type locality: South Africa, Western Cape, Cape of Good Hope [as “cap de Bonne-Espérance”].
Reaumuria lalandii Robineau-Desvoidy, 1830: 80. Type(s), unspecified sex (MNHN or lost). Type locality: South Africa, Western Cape, Cape of Good Hope [as “cap de Bonne-Espérance”].
Note: Gonia fasciata Wiedemann, 1819 was described from an unspecified number of females, but certainly more than one because Wiedemann (1930: 344) later wrote, “In Westermann’s und meiner Sammlung”. Hence, the original type series is interpreted as consisting of syntypes.
madagascariensis Villeneuve, 1915.—Afrotropical: Madagascar.
Pseudogonia madagascariensis Villeneuve, 1915b: 192. Lectotype female (CNC), by fixation of Cooper and O’Hara (1996: 66) (data on “Holotype ♀” from Tananarive in CNC is regarded as a lectotype fixation). Type locality: Madagascar, Antananarivo, Antananarivo [as “Tananarive”].
Note: There are eight paralectotypes, most or all of which females, of Pseudogonia madagascariensis Villeneuve, 1915 in NHMW (examined by JEOH).
rufifrons (Wiedemann, 1830).—Afrotropical: “widespread Afrotrop. Reg.” (Crosskey 1980b: 875), including Cape Verde, Nigeria, South Africa, Tanzania, U.A. Emirates, Yemen. Palaearctic: C. Asia, Europe (all except British Is., Scand.), Japan, Kazakhstan, Korea (S. Korea), M. East (Israel), Mongolia, Pal. China, Russia (W. Russia, W. Siberia, S. Far East), Transcaucasia. Oriental: India, Indonesia, Malaysia, Myanmar, Orien. China, Pakistan, Philippines, Ryukyu Is., Taiwan, Thailand. Australasian: Australia, Hawaii, Melanesia, N. Australasian.
Latreillia lalandii Robineau-Desvoidy, 1830: l06 (junior secondary homonym of Reaumuria lalandii Robineau-Desvoidy, 1830). Type(s), unspecified sex (MNHN or lost). Type locality: South Africa, Western Cape, Cape of Good Hope [as “cap de Bonne-Espérance”].
Tachina rufifrons Wiedemann, 1830: 318. Lectotype female (ZMUC), by fixation of Crosskey (1966a: 677) (examination of “Holotype ♀” from China in ZMUC is regarded as a lectotype fixation). Type locality: China.
Gonia cinerascens Rondani, 1859: 34. Syntypes, unspecified number and including at least 1 male (MZF, Herting 1969: 192; 1 male and 7 females in MZF [examined by PC]). Type locality: Italy, hills near Parma.
Gonia munroi Curran, 1927d: 339. Holotype male (BMNH). Type locality: Tanzania.
Gonia ritchiei Cuthbertson & Munro, 1941: 109. Nomen nudum.
Note: The relative priority of Reaumuria lalandii Robineau-Desvoidy, 1830 and Latreillia lalandii Robineau-Desvoidy, 1830, when both are placed in the same genus, was established by Crosskey (1980b: 875), as the First Reviser (Article 24.2.2 of the Code, ICZN 1999). Since the latter name was given junior homonym status, it cannot replace Tachina rufifrons Wiedemann, 1830 as the valid name of the species even though it was published first; see dating of Robineau-Desvoidy (1830) and Wiedemann (1830) in References.
Gonia cinerascens Rondani, 1859 was probably described from both sexes but the original description only made specific mention of the male. Crosskey (1976: 244) reported three male and four female syntypes in MZF. An examination of the MZF holdings by PC discovered another syntype and a change to the reported sexes. The type series was found to consist of one male and seven females. The single male is Gonia picea (Robineau-Desvoidy, 1830) and the seven females conform to the present concept of Gonia cinerascens Rondani. When treated as a valid name, G. cinerascens has also been called Isomera cinerascens (Rondani) in the literature.
Gonia munroiCurran (1927d: 339) was “Described from 5♂♂, 8♀♀, from Tanganyika and South Africa. The type male and female are from Tanganyika ...”. We recognize the male from Tanzania as a designated holotype whereas Arnaud and Owen (1981: 209) treated all specimens as syntypes, writing “syntypes, 5 males and 8 females”.
suspecta Villeneuve, 1915.—Afrotropical: Madagascar.
Pseudogonia suspecta Villeneuve, 1915b: 192 (as “P. suspecta (n. sp.? vel n. var.?)”). Syntypes, 3 specimens of uncertain sex [“Par l’absence de soies orbitaires et surtout par la longueur des antennes, ils semblent bien être des ♂; néanmoins les griffes sont courtes comme chez les ♀.”] (NHMW, not located by JEOH). Type locality: Madagascar.
Genus RAMONELLA Kugler, 1980
RAMONA Kugler, 1980a: 40 (junior homonym of Ramona Casey, 1886). Type species: Ramona mesnili Kugler, 1980, by original designation.
RAMONELLA Kugler, 1980b: 67 (nomen novum for Ramona Kugler, 1980).
mesnili (Kugler, 1980).—Afrotropical: Yemen. Palaearctic: Europe (Turkey), M. East (Israel), N. Africa (Canary Is.).
Ramona mesnili Kugler, 1980a: 41. Holotype male (TAU). Type locality: Israel, Negev, Ramon.
Genus RHYNCHOGONIA Brauer & Bergenstamm, 1893
RHYNCHOGONIA Brauer & Bergenstamm, 1893: 37, 104 [also 1893: 125, 192]. Type species: Rhynchogonia algerica Brauer & Bergenstamm, 1893, by monotypy.
algerica Brauer & Bergenstamm, 1893.—Afrotropical: U.A. Emirates. Palaearctic: C. Asia, M. East (Israel), N. Africa (NW. Africa).
Rhynchogonia algerica Brauer & Bergenstamm, 1893: 105 [also 1893: 193]. Type(s), female (1 female in NHMW according to Herting 1974b: 135, not located by JEOH). Type locality: “Afrika” (Algeria, Biskra according to Herting 1974b: 135).
Genus SCHEMBRIA Rondani, 1861
SCHEMBRIA Rondani, 1861b: 110. Type species: Schembria meridionalis Rondani, 1861, by monotypy [Palaearctic].
Note: Schembria Rondani, 1861 was first recognized from the Afrotropical Region by Crosskey (1984: 201, 287) based on an undescribed species from South Africa. That species was subsequently described by Barraclough (1991: 135) as Schembria eldana.
eldana Barraclough, 1991.—Afrotropical: South Africa.
Schembria eldana Barraclough, 1991: 135. Holotype male (NMDA). Type locality: South Africa, KwaZulu-Natal, Lower Tugela River, Tongaat (29°35′S 31°08′E), Wewe Sugar Estate.
Genus SIMOMA Aldrich, 1926
SIMOMA Aldrich, 1926b: 20. Type species: Simoma grahami Aldrich, 1926, by original designation. New record.
grahami Aldrich, 1926.—Afrotropical: Namibia (new record, MZUR [PC]). Palaearctic: Japan, M. East (Israel), Pal. China. Oriental: India, Malaysia, Orien. China, Vietnam.
Simoma grahami Aldrich, 1926b: 21. Holotype male (USNM). Type locality: China, Sichuan, Suifu.
Note: Simoma grahami Aldrich, 1926 is newly recorded from the Afrotropical Region. O’Hara et al. (2009: 117), in a note about Simoma grahami, wrote: “This species may have been recorded from Japan in error (e.g., Crosskey 1976: 253, Herting 1984: 73)”. Simoma grahami has since been recorded from Honshu (Japan) by Shima (2014: 861) and PC examined a male in IRSNB from Tokyo (collected by Edme Gallois on 6 June 1909).
Genus STIREMANIA Cerretti & O’Hara, gen. n.
STIREMANIA Cerretti & O’Hara, gen. n. Type species: Stiremania karoo Cerretti and O’Hara sp. n., by present designation.
Note: This new genus and the two new species below are described in the New Taxa of Afrotropical Tachinidae section.
karoo Cerretti & O’Hara, sp. n.—Afrotropical: South Africa.
Stiremania karoo Cerretti & O’Hara, sp. n. Holotype male (MZUR). Type locality: South Africa, Western Cape, Gamkaskloof (Die Hel), 336m (33°22′5.90″S 21°37′19.43″E).
robusta Cerretti & O’Hara, sp. n.—Afrotropical: South Africa.
Stiremania robusta Cerretti & O’Hara, sp. n. Holotype male (NMDA). Type locality: South Africa, Eastern Cape, Willowmore.
Genus STURMIA Robineau-Desvoidy, 1830
STURMIA Robineau-Desvoidy, 1830: 171. Type species: Sturmia vanessae Robineau-Desvoidy, 1830 (= Tachina bella Meigen, 1824), by subsequent designation of Robineau-Desvoidy (1863a: 888) (earlier type fixations set aside by ICZN 2012: 242; see Evenhuis and Thompson 1990: 238 and O’Hara and Evenhuis 2011: 61) [Palaearctic].
POLYCHNOMYIA Bischof, 1904: 85. Type species: Polychnomyia flavohalterata Bischof, 1904 (= Tachina convergens Wiedemann, 1824), by monotypy.
VERBEKEIA Mesnil, 1959: 5. Type species: Verbekeia lindneri Mesnil, 1959, by monotypy.
bellina Mesnil, 1944.—Afrotropical: Madagascar.
Sturmia bellina Mesnil, 1944: 10. Holotype male (MNHN). Type locality: Madagascar, Toliara, Bekily.
convergens (Wiedemann, 1824).—Afrotropical: Ethiopia, Kenya, Malawi, Nigeria, Sierra Leone, South Africa, Tanzania, Uganda, Zambia, Zimbabwe. Oriental: India, Sri Lanka. Australasian: Australia, N. Australasian.
Tachina convergens Wiedemann, 1824: 43. Lectotype female (ZMUC), by designation of Crosskey (1963: 78). Type locality: “India orient.” (i.e., “East Indies”; interpreted as India by Crosskey 1963: 78 and Crosskey 1976: 242).
Polychnomyia flavohalterata Bischof, 1904: 86. Type(s), male (1 male in NHMW). Type locality: South Africa, Eastern Cape, Algoa Bay.
Sturmia completa Curran, 1927f: 119. Holotype male (SANC). Type locality: South Africa, Mpumalanga, White River [ca. 25°20′S 31°1′E].
Tachina bella of authors, not Meigen, 1824. Misidentification (Crosskey 1980b: 874).
Note: Polychnomyia flavohalterata Bischof, 1904 was described from one or more males. There are two specimens in NHMW, one male and one female, with label data corresponding to that published for the type(s) of P. flavohalterata and both labelled as “Typ.” by Villeneuve (there is no Bischof det. label on either of these specimens). Since the original description mentioned only the male sex, only the male is a name-bearing type. This male also bears a Crosskey holotype label dated 1970, but Bischof did not give the number of specimens upon which his description was based and therefore this “holotype” is regarded as a syntype.
lindneri (Mesnil, 1959).—Afrotropical: D.R. Congo, Nigeria, Tanzania, Uganda.
Verbekeia lindneri Mesnil, 1959: 5. Holotype male (SMNS). Type locality: Tanzania, Kware [ca. 3°17′S 37°9′E].
profana (Karsch, 1888).—Afrotropical: “Ost-Afrika” [East Africa]. Comb. n.
Degeeria profana Karsch, 1888: 376. Holotype male [not female as published] (ZMHB). Type locality: “Ost-Afrika” [East Africa].
Note: Degeeria profana Karsch, 1888 was treated as an unplaced species of “Goniinae” [= Exoristinae] by Crosskey (1980b: 881) but is moved here to Sturmia Robineau-Desvoidy, 1830 based on examination of the holotype.
rasa (Mesnil, 1959).—Afrotropical: Tanzania.
Pimelimyia rasa Mesnil, 1959: 8. Holotype male (SMNS). Type locality: Tanzania, “Ngaruka” [probably Engaruka, ca. 3°0′S 35°58′E].
rasella (Mesnil, 1970).—Afrotropical: Madagascar.
Pimelimyia rasella Mesnil, 1970b: 100. Holotype male (MNHN). Type locality: Madagascar, Toliara, Sakaraha.
velutina Mesnil, 1944.—Afrotropical: Madagascar.
Sturmia velutina Mesnil, 1944: 11. Holotype male (MNHN). Type locality: Madagascar.
Unplaced species of Goniini
clarior Villeneuve, 1943.—Afrotropical: Zimbabwe.
Sturmia russata clarior Villeneuve, 1943a: 38. Holotype male (not located). Type locality: southern Zimbabwe.
inimica Hesse, 1934.—Afrotropical: South Africa.
Sturmia inimica Hesse, 1934: 428. Holotype female (SAMC). Type locality: South Africa, Western Cape, Somerset West.
rufiventris Curran, 1927.—Afrotropical: D.R. Congo. Comb. n.
Ceromasia rufiventris Curran, 1927c: 7. Holotype female (AMNH). Type locality: D.R. Congo, Orientale, Kisangani [as “Stanleyville”].
Note: Ceromasia rufiventris Curran, 1927 was placed in Eurysthaea Robineau-Desvoidy, 1863 by Crosskey (1980b: 878) as a new combination. The female holotype was examined by PC and could not be placed to genus within the Goniini but does not belong to Eurysthaea.
vocalis Villeneuve, 1943.—Afrotropical: D.R. Congo.
Sturmia vocalis Villeneuve, 1943a: 36. Syntypes, 1 male and 1 female (not located). Type localities: D.R. Congo, Orientale, Kisangani [as “Stanleyville”] and Nord-Kivu, “Lessewoud” [assumed to be Lesse at ca. 0°45′N 29°46′E].