The evolution of the male endophallus in the subgenus Circinatus based on the most parsimonious tree obtained with the SW analysis. Black: endophallus type I; gray: endophallus type II; white: endophallus type III; black-white strips show the intermediate status between endophallus types I and III. The species with an unknown endophallus are marked with an underline.

  
 
  Part of: Shi H, Liang H (2015) The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini). ZooKeys 536: 1-92. https://doi.org/10.3897/zookeys.536.5982