Single most parsimonious tree (1392 steps) based on 53 morphological characters. Bootstrap (B) values over 50% are noted above the corresponding branches, respectively. Branch support was calculated by bootstrap with 10000 replicates. Character states are shown above branches.

  Part of: Keresztes L, Kappert J, Henning M, Török E (2021) Helen’s twins in the Balkans: discovery of two new Paraptychoptera Tonnoir, 1919 species closely related to P. helena Peus, 1958, with systematic revision of the “lacustris” group (Diptera, Ptychopteridae). ZooKeys 1071: 63-81.