Upper dentition of L. dekeyseri A, C (LDM 3185) and L. mordax B, D (ALP 6149). A, B Moderate inner lobe in the first upper premolar (P3) of L. dekeyseri A contrasting with the lingual lobe of P3 absent or very reduced in L. mordax B (similar condition observed in L. inexpectata) C, D metastyles of M1 and M2 reduced or absent in dekeyseri C contrasting with the metastyles well developed and distinct in L. inexpectata and L. mordax D.