Tree topology obtained by Maximum Likelihood in RAxML using a modified version of Lopardo et al. (2011) and Feng et al. (2019) plus the four symphytognathid species from our study (in red). Numbers at nodes indicate bootstrap support. Note the long branch of Anapistula and its position within Anapidae; and the high support of Crassignatha and Patu in the Symphytognathidae. Molecular vouchers used for previous “symphytognathoid” studies (Lopardo et al. 2011; Lopardo and Hormiga 2015) identified to genus level by L. Lopardo (pers. comm.) as follows: ■ Crassignatha (apparently conspecific with C. seeliam); ◆Patu; and ▲Symphytognatha.