Riotintobolus mangatsiaka sp. nov., male and female (FMMC 5413), SEM A left gnathal lobe of mandible, mesal view B left female vulva, lateral view. Abbreviations: av = anterior orientated plate; 3iT = 3-combed inner tooth; eT ? external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamellae; pv = posterior orientated plate. Scale bars as indicated.

  Part of: Wesener T (2020) Ecotone shifts in southern Madagascar: first barcoding data and six new species of the endemic millipede genus Riotintobolus (Spirobolida, Pachybolidae). ZooKeys 953: 1-29.