Riotintobolus mangatsiaka sp. nov., paratypes (ZFMK MYR938) A living female B living male in defensive position C multi-layer photograph, male anterior body, lateral view D multi-layer photograph, male leg pairs 1–7, ventral view E multi-layer photograph, male posterior body end with telson F anterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, posterior view I right posterior gonopod, anterior view. Abbreviations: av = paraprocts; Co = collum (ring 1); Cx = coxite; Md = basal joints of mandible; Pe = penis; Pre = epiproct; St = sternite; sub = hypoproct; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ‘flag’; numbers refer to leg pair number. Scale bars: 1 mm.

  Part of: Wesener T (2020) Ecotone shifts in southern Madagascar: first barcoding data and six new species of the endemic millipede genus Riotintobolus (Spirobolida, Pachybolidae). ZooKeys 953: 1-29.