Relationships of Riotintobolus species. The evolutionary history of Riotintobolus was inferred by using the Maximum Likelihood method based on the Tamura-Nei model. The tree with the highest log likelihood (-5174.8354) is shown. The tree is drawn to scale, with branch lengths measured in the number of substitutions per site. Yellow box marks the genus Riotintobolus, green colour bars mark the new species described here. For specimen information, see Table 1. Numbers refer to bootstrap values, values < 50% not shown.