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  • Article title
  • Abstract
  • Keywords
  • Introduction
  • Materials and methods
    • Morphology
    • Phylogeny
    • Biogeographic origin of the P. lacrimosus species group
  • Results
    • Phylogeny and biogeography
    • Hyperdistal subarticular tubercles and phalange morphology
    • Systematic accounts
      • Pristimantis lacrimosus species group
  • Discussion
    • Subarticular tubercles and phalange morphology in Pristimantis
    • The Pristimantis lacrimosus species group originated in the Chocoan forests
  • Acknowledgments
  • References
  • Appendix 1
    • Examined specimens
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