Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species

1, 2 Pontohoratia vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 1 holotype 2 paratype 3 P. pichkhaiai sp. nov., Samegrelo, Mukhuri, Shisha Spring, holotype 4 Hausdorfenia pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring, holotype 5 H. shareula sp. nov., Racha, Nikortsmintha, Tsivtskala 2 Spring near power station in the Shareula valley, holotype 6 P. mapeli sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring, holotype 7 H. pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring; aberrant specimen 8, 9 P. vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 8 morphotype 9 live specimen with marked carbonate incrustations. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.