Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species

Distribution map of genera Pontohoratia Vinarski, Palatov & Glöer, 2014; Motsametia Vinarski, Palatov & Glöer, 2014 and Hausdorfenia gen. nov. 1 P. smyri Vinarski, Palatov & Glöer, 2014 (black dot) 2–4 P. birsteini (Starobogatov, 1962) (grey dots) 6 M. borutzkii (Shadin, 1932) (purple dot) 10 H. shareula sp. nov. (dark red dot) 12–15 H. hauffeniaformis sp. nov. (red dots) 27 P. pichkhaiai sp. nov. (light blue dot) 28 P. mapeli sp. nov. (medium blue dot) 30–31 P. vinarskii sp. nov. (dark blue dot).