Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species

Distribution map of Imeretiopsis gen. nov.: (magenta dots) and Caucasopsis gen. nov. (blue dots) 6 I. cameroni sp. nov. and I. iazoni sp. nov. 7 Imeretiopsis prometheus sp. nov. 8 I. nakeralaensis sp. nov. 9 I. gorgoleti sp. nov. 0 Caucasopsis orientalis (Starobogatov, 1962), C. subovata (Starobogatov, 1962) and C. pulcherrima (Starobogatov, 1962) 3 C. shadini (Starobogatov, 1962) 4 C. aculeus (Starobogatov, 1962) and C. schakuranica (Starobogatov, 1962) 22, 23 C. egrisi sp. nov. 24 C. olsavskyi sp. nov. 25, 30, 31 C. letsurtsume sp. nov.