Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species

1–5 Kartvelobia sinuata sp. nov., Imereti, Satsiskvilo, Turchusmtha 1 holotype 2–5 paratypes 6 K. sinuata sp. nov., Imereti, Turchusmtha, spring of Okatse above Kinchkha waterfall 7, 8 K. cf. sinuata, Samegrelo, Pirveli Balda, Motena Cave 9–11 K. cf. sinuata, Samegrelo, Pirveli Balda, spring in village 12 K. sinuata sp. nov., Imereti, Nakhriduri, spring cave Turchu Gamosadivari 13 K. sinuata sp. nov., Imereti, Nakhriduri spring at Turchu Gamosadivari Basin 14, 15 K. sinuata sp. nov., Imereti, Nakhriduri side spring at Turchu Gamosadivari Basin. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.