Description of 47 new species of the New Caledonian endemic caddisfly genus Agmina Ward & Schefter (Trichoptera, Ecnomidae)

Abstract New Caledonia has a rich Trichoptera fauna with over 200 known species, most of them endemic. The total diversity has been estimated as high as 300 to 600 species. The endemic genus Agmina Ward & Schefter (Ecnomidae, Trichoptera) includes 28 described species. Based on male genitalia morphology and previously published molecular data another 47 new species in the genus are described, namely Agmina tuberosasp. nov., A. semiovalesp. nov., A. rochetasp. nov., A. tenuisasp. nov., A. multidentatasp. nov., A. cornutasp. nov., A. sagittatasp. nov., A. circulatasp. nov., A. digitatasp. nov., A. longispinasp. nov., A. magnahamatasp. nov., A. longicordatasp. nov., A. campanulasp. nov., A. semicampanulasp. nov., A. cuniculasp. nov., A. cerritulasp. nov., A. monstrosasp. nov., A. rectangulatasp. nov., A. chelasp. nov., A. piscariasp. nov., A. amplexasp. nov., A. caraffa, sp. nov., A. rostratasp. nov., A. dathioensissp. nov., A. rougensissp. nov., A. viklundisp. nov., A. latasp. nov., A. falxsp. nov., A. guttulasp. nov., A. amieuensissp. nov., A. spinasp. nov., A. complexasp. nov., A. dognyensissp. nov., A. manasp. nov., A. anterohamatasp. nov., A. curvatacuasp. nov., A. recurvatasp. nov., A. taoensissp. nov., A. triangulatasp. nov., A. bleuensissp. nov., A. touhoensissp. nov., A. wardisp. nov., A. parallelasp. nov., A. christinaesp. nov., A. brevissp. nov., A. ninguanasp. nov., and A. scopulasp. nov. Additionally, new records are provide for the species A. acula Ward, 2003, A. artarima Ward & Schefter, 2000, A. berada Ward & Schefter, 2000, A. bimaculata Ward & Schefter, 2000, A. cheirella Ward, 2003, A. comata Ward, 2003, A. diriwi Ward & Schefter, 2000, A. hamata Ward & Schefter, 2000, A. hastata Ward & Schefter, 2000, A. hirta Ward & Schefter, 2000, A. jepiva Ward & Schefter, 2000, A. joycei Ward & Schefter, 2000, A. kapiwa Ward & Schefter, 2000, A. kara Ward & Schefter, 2000, A. mariae Ward & Schefter, 2000, A. nodosa Ward, 2003, A. panda Ward & Schefter, 2000, A. padi Ward & Schefter, 2000, A. parie Ward & Schefter, 2000, A. rhara Ward & Schefter, 2000, A. urugi Ward & Schefter, 2000, and A. vuegi Ward & Schefter, 2000. With a total of 75 described species Agmina is one of the largest animal radiations in New Caledonia. Nothing is known about the early stages of any of the species in this genus.


introduction
New Caledonia in the southwest Pacific is the smallest of the original biodiversity hotspots (Myers et al. 2000) and has a rich both terrestrial and freshwater insect fauna (e.g., Balke et al. 2007;Espeland and Johanson 2010a, b;Papadopoulou et al. 2013;Gibbs and Lees 2014). The islands have been shown to be especially rich in caddisflies (Trichoptera) with almost 250 described species and an estimated total of 300 to 600 species (Espeland and Johanson 2010a;Johanson and Wells 2019). The caddisfly genus Agmina Ward & Schefter, 2000 (Ecnomidae) currently contains 28 species all endemic to New Caledonia. Approximately 40% of these species were described based on single specimens and it has been predicted that the total number of species is probably much higher Johanson and Wells 2019;Wells et al. 2019). Support for this comes from a phylogenetic study showing that there are at least 47 undescribed species in the genus, making it one of the largest known animal radiations on New Caledonia (Espeland and Johanson 2010), now possibly surpassed only by eumolpine leaf beetles (Papadopoulou et al. 2013). The study by Espeland and Johanson (2010) indicated that Agmina split from its closest relatives around 36 mya just after the island reappeared from being submerged for much of the Palaeocene and first half of the Eocene (e.g., Paris 1981;Aichison et al. 1998;Crawford et al. 2003;Pelletier 2007;Cluzel et al. 2012). The earliest radiation was dated to the early Miocene. More than half the species appear to be adapted to the nutrient poor, but toxic, nickel-rich ultramafic substrate currently found on approximately one third of the island (Espeland and Johanson 2010), which generally has been shown to have a poor aquatic macroinvertebrate diversity compared to other substrates (Mary 2002).
Here we describe 47 new species in the genus Agmina increasing the number of known species by 270%, to 75 species. Several of the new species are based on singletons shown to be good species in the molecular phylogeny of Espeland and Johanson (2010). With this report, 286 species of caddisflies are now known from New Caledonia.

Materials and methods
The material used in this study was collected in Malaise traps and light traps on the New Caledonian Grande Terre during three expeditions between 2001 and 2006. The Malaise traps were set in place for approx. two weeks at a time, the light traps were operated overnight only. All material was sampled directly into 80% alcohol and transported to the laboratory at the Swedish Museum of Natural History (NHRS) for sorting and determination. The determinations were carried out using the information in Ward and Schefter (2000) and . Material of all Agmina species in the samples was included in a phylogenetic analysis (Espeland and Johanson 2010a) which resulted in a tree with seven monophyletic clades including 22 previously described species and 47 undescribed species. The seven clades are used below to group the species according to the phylogenetic position rather than morphological similarities, and diagnosis of the species groups are therefore not included. Extraction of DNA from the specimens was done from the individual abdomens, which were macerated during that process. The abdomens were dehydrated in absolute alcohol and temporarily mounted in Euparal on a microscope slide before examination and drawing. All drawings were produced in pencil on plain white A4 paper sheets using a drawing tube mounted on a Leitz Ortholux II. After the drawings were completed the abdomens were returned to the alcohol vial with the rest of the animal. Each pencil illustration was digitised in a scanner at low resolution and thereafter used as a background layer in Adobe Photoshop 8.0. The illustrations were completed after being re-drawn on a new layer using a Wacom drawing pad before the background layer was deleted. The nomenclature applied to the genitalic morphology follows that of Johanson (2017). Specimens in this study are deposited in the following repositories: Diagnosis. Agmina tuberosa sp. nov. resembles many other Agmina species in having large, oval superior appendage, but is distinguished from other species by the large inferior appendages, each with a long dorsal and short ventral branch in lateral view, and the sternal processes reaching to half length of the inferior appendages. The genitalia are similar to those of A. semiovale sp. nov. from which it is distinguished by the presence of a row of teeth-like megasetae on the mesal margin of each paramere in dorsal view. Etymology. Tuberosa, from Latin, meaning potato, named for the superior appendage being potato-shaped in lateral view.
Description. Genitalia: In lateral view, segment IX widely rounded anteriorly, apex located dorsally; in ventral view anteriorly widely U-shaped. Sternal processes, lateral view, with apex not exceeding posterior apex of tergum X, narrowing along their length, curved ventro-posteriorly at mid-length; in ventral view, slender, straight, slightly diverging along their length. Tergum X smoothly convex dorsally, in lateral view longer than high; in dorsal view, mesally separate, axe-shaped with straight inner margins. Parameres robust, starting before tergum X, long, narrow; in lateral view running parallel with dorsal margin of segment IX, X, and superior appendage, ending before apex of superior appendage; in dorsal view, separated and narrowing along their length, each with truncate apex, inner margin with row of megasetae. Superior appendage, in lateral view, large, oval, with posterad spine-like mesal process present on ventromesal margin exceeding the main branch posteriorly; in dorsal view uniformly narrow, running almost parallel, slightly curving inwards towards blunt apex. Mesal processes straight, orientated slightly mesally. Inferior appendage with posterad orientated long dorsal branch slightly dorsally curving at acute apex; dorsal branch widely separated from short, triangular, ventral branch; in ventral view large, wide, oval, with posterad orientated dorsal branches, ventral branch forming central lobe. Phallus, in lateral view as long as segment IX, slender and slightly curving downwards; in ventral view uniformly tapering along its length.
Additional information. This species was referred to as "sp. 26" in Espeland and Johanson (2010). Diagnosis. Agmina semiovale sp. nov. resembles many other Agmina species in having a large semi-oval shaped superior appendage in lateral view. The species is distinguished from the other species by the presence of very large parameres that partly exceeds the superior appendages dorsally as seen in lateral view, and in dorsal view forming an oval basis and two broad posterad branches. The genitalia are similar to those of A. tuberosa sp. nov. from which it is distinguished by the absence of a row of teeth-like megasetae on the mesal margin of each paramere.
Etymology. Semiovale, from Latin, meaning half oval, named for the shape of the superior appendage in lateral view.
Description. Genitalia: Segment IX damaged, with sternal processes missing. Tergum X sub-rectangular; in dorsal view, mesally separate, more than two times longer than wide. Parameres robust, starting before tergum X, in lateral view long, wide at base, first half narrowing towards apex, apical half slightly widening, ventral margin slightly concave; in dorsal view, fused at base, apical 2/3 rds separated, each with truncated apex, lateral margins convex, mesal margins slightly concave. Superior appendages, in lateral view, longer than tergum X, longer than wide, narrowing along its length, dorsal margin straight, ventral margin convex, with posterad spine-like mesal process present on ventromesal margin not exceeding posteriorly the main branch; in dorsal view slightly narrowing towards apex, running almost parallel, almost straight with blunt apex slightly curving inwards. Mesal processes straight, orientated slightly mesally. Inferior appendages missing due to damage. Phallus, in lateral view approx. as long as segment IX, three times longer than wide; in ventral view proximal half tapering along its length, apical half straight and tapering at apex.
Additional information. This species was referred to as "sp. 57" in Espeland and Johanson (2010).  very long and curved mesal process reaching as far posteriorly and above the apex of the superior appendages. It particularly resembles A. nodosa  but in A. nodosa the sternal processes are simple while in A. rocheta sp. nov. they are bifurcated.
Etymology. Rocheta (noun, feminine), from Latin, meaning rocket. Named for the rocket-shaped phallus in ventral view.  Agmina padi , particularly in the shape of the inferior appendages in lateral view. Agmina tenuisa sp. nov. is distinguished from A. padi by the more slender superior appendages in lateral view, the ventral branch of the parameres exceeding below the superior appendages, and in ventral view the plate of inferior appendages has no central posterior plate.
Etymology. Tenuisa, from Latin, meaning slender. Named for the slender superior appendages as seen in lateral view.
Description. Genitalia: In lateral view, segment IX triangular anteriorly, apex located medially; in ventral view anteriorly U-shaped. Sternal processes, lateral view, with each apex not exceeding posterior margin of tergum X, short, triangular; in ventral view, short, not reaching mid-length of inferior appendage, slightly diverging along their length, apices slightly curving mesad. Tergum X small, in lateral view around half the length of inferior appendage, dorsal margin straight, ventral margin slightly convex; in dorsal view, mesally separate, semi-trapezoid, outer margins straight, inner margin slightly convex. Parameres starting at anterior margin of tergum X; in lateral view, long, slender, widely U-shaped, with posterior end slightly diverging, apex pointing posteroventrally; in dorsal view, separated, narrow, posterior 1/3 directed posteromesad, each with short, thick, setae at apex. Superior appendages, in lateral view, slender, slightly curving upwards, more than twice as long as tergum X, with spine-like process at apex, mesal process spine-like straight, slightly curving downwards at base, exceeding posteriorly the main branch; in dorsal view around four times longer than wide, almost parallel, each slightly curving inwards toward apex with thin posteromesad orientated process with long thick seta. Mesal processes long, slender, gently curving mesad, crossing each other near apex. Inferior appendages, in lateral view, with posterodorsally orientated claw-shaped dorsal branch, ventral branch short, claw-shaped, orientated posterodorsally, originating at approx. mid-length on ventral margin of dorsal branch; in ventral view small, oval at base with slender, posteriorly orientated dorsal branches, slightly converging towards apex, ventral branches directed posterolaterally, each with claw-shaped apex. Phallus, in lateral view as long as segment IX, slightly sigmoid; in ventral view anterior half distinctly wider than posterior half.
Additional information. This species was referred to as "sp. 12" in Espeland and Johanson (2010). ages forming rhomboid plates in lateral view; and tergum X forms a transverse bridge. It resembles A. cornuta sp. nov. in the shape of tergum X, but in A. multidentata sp. nov. the inferior appendages are higher in lateral view and the superior appendages are more prolonged posteriorly compared to those in A. cornuta sp. nov.
Etymology. Multidentata, from Latin, meaning many teeth. Named for the ventral and mesal margins of the superior appendages with many teeth as seen in lateral view.
Description. Genitalia: In lateral view, segment IX triangular anteriorly, apex located mediodorsally; in ventral view anteriorly semi-circular. Sternal processes, lateral view, with each rounded apex not exceeding posterior apex of tergum X, triangular; in ventral view, broad, reaching mid-length of inferior appendage, slightly diverging, apices curving mesad. Tergum X small, in lateral view less than half the length of superior appendage, slightly longer than high, inner and outer margins almost straight, slightly diverging anteriorly, anterior margin concave; in dorsal view, mesally connected, axeshaped. Parameres complex; in lateral view with tubular dorsal lobe running parallel with dorsal margin of tergum X and superior appendage, ending before mid-length of superior appendage; shorter, narrower, ventral lobe orientated posteroventrally; in dorsal view, dorsal lobes separated and narrowing along their length, not reaching mid-length of superior appendage, directed posteromesad, each with truncate apex. Superior appendages, in lateral view, more than twice as long as tergum X, tapering along its length with rounded base, apex semi-acute with stout megaseta orientated ventrad mesal process absent; in dorsal view uniformly narrow, running almost parallel, slightly curving inwards towards apex with stout megaseta directed mesad, inner margin dentate. Mesal processes absent. Inferior appendages, in lateral view, with posterad orientated triangular dorsal branch with acute apex; dorsal branch present as short, obtuse process at posteroventral margin in ventral view convex at base, gradually widening until mid-length, then tapering towards apex, dorsal branches orientated posteriorly, claw-shaped, with apices directed posteromesad, ventral branch barely visible as short very obtuse central lobe. Phallus, in lateral view as long as segment IX, almost straight with slightly concave margins in ventral view anterior half not distinctly wider than posterior half.
Additional information. This species was referred to as "sp. 10" in Espeland and Johanson (2010 Diagnosis. Agmina cornuta sp. nov. is distinguished from the other Agmina species by the presence of a pair of almost uniformly oval superior appendages in lateral view, and a pair of branches of the inferior appendages which appear beak-like in lateral view and are approx. as long as the superior appendages. It resembles A. multidentata sp. nov. in the shape of tergum X, but in A. cornuta sp. nov. the inferior appendages are lower in lateral view and the superior appendages are rounder posteriorly compared to those in A. multidentata sp. nov. Etymology. Cornuta, from Latin, meaning horned. Named for the shape of the inferior appendage in lateral view, which has two long, horn-shaped, dorsal branches. Measurements. Fore wing length 3.2-4.1 mm (N = 4). Total length of genitalia: 0.5 mm.
Description. Genitalia: In lateral view, segment IX semi-triangular anteriorly, apex located dorsally; in ventral view anteriorly widely U-shaped. Sternal processes, lateral view, with each apex, not exceeding posterior apex of tergum X, very short, triangular; in ventral view, short, slender, not reaching mid-length of inferior appendage, each with apex slightly curving mesad. Tergum X small, in lateral view approx. half the length of superior appendage, outer margin irregularly convex, slightly converging towards posterior with straight inner margin; in dorsal view, mesally connected, axe-shaped with inner margins forming a right triangle. Parameres starting at tergum X, in lateral view semi-circular, each with narrow protrusion orientated posteroventrad with blunt apex, approx. half the length of the main parameres; in dorsal view, separated, slightly diverging towards apices, outer margins straight for first 2/3, then abruptly curving inwards at obtuse angle, inner margin irregularly lobed. Superior appendages, in lateral view, approx. twice as long as tergum X, semiovoid with truncated, irregularly dentate apex, mesal process absent; in dorsal view almost parallel, each gently widening to mid-length, then inner margin abruptly widening, with apical half irregularly dentate. Mesal processes absent. Inferior appendages, in lateral view, with posterad orientated long, knife-shaped dorsal branch; ventral branch present as short, obtuse process at posteroventral margin; in ventral view convex at base, gradually widening for first 1/3, then abruptly narrowing towards apex, long, narrow, slightly tapering dorsal branches orientated posteriorly, slightly curving inwards, apices pointed, ventral branch indiscernible. Phallus, in lateral view shorter than segment IX, irregular; in ventral view posterior half slightly wider than anterior half.
Additional information. This species was referred to as "sp. 53" in Espeland and Johanson (2010).

Species group 2, sagittata group
This group includes the following species: Agmina sagittata sp. nov., A. circulata sp. nov., A. jepiva , A. digitata sp. nov., A. longispina sp. nov., A. urugi Ward & Schefter, 2000, A. magnahamata sp. nov., A. longicordata sp. nov., A. campanula sp. nov., A. semicampanula sp. nov., A. cunicula sp. nov., A. panda Ward & Schefter, 2000 Diagnosis. Agmina sagittata sp. nov. is distinguished from the other Agmina species in the genitalia by the presence of a pair of inferior appendages having a ventral part that, in lateral view, reaches as far posteriorly as the sternal processes, and the apex of the sternal processes and inferior appendages are separated by a rectangular incision. It resembles A. circulata sp. nov. in the overall similarity of the genitalia, but A. circulata sp. nov. is larger than A. sagittata sp. nov., and in the genitalia A. sagittata sp. nov. has superior appendages that are more rounded in lateral view, shorter sternal processes in relation to the inferior appendages, and the sternal processes of A. sagittata sp. nov. is slightly wider and straighter than those of A. circulata sp. nov.
Etymology. Sagittata, from Latin, meaning arrow-shaped. Named for the inferior appendages being arrow-shaped in ventral view.
Description. Genitalia: Total length 0.4 mm. In lateral view, segment IX semitriangular anteriorly, apex located dorsally; in ventral view anteriorly obround. Sternal processes, in lateral view, with each apex exceeding the length of tergum X, elongated narrowly triangular with semi-acute apex, dorsal margin almost straight, ventral margin gently concave; in ventral view, longer than inferior appendage, narrowing along their length, apex acute. Tergum X in lateral view irregularly quadrilateral, slightly longer than superior appendage, longer than wide, tapering along its length; in dorsal view, mesally separate, axe-shaped, inner margins forming triangle. Parameres start-  ing before tergum X, in lateral view long, slowly widening along 2/3 of its length, then narrow, apex slender, multifurcated, ending before apex of superior appendage; in dorsal view, separated, long, slender, straight, apex pipe wrench-shaped. Superior appendages, in lateral view, shorter than tergum X, semi-trapezoid; in dorsal view stout, curving inwards, with rounded outer margin; mesal process, stout claw-like, directed mesad. Inferior appendages, in lateral view, with posterad orientated short, rectangular dorsal branch; ventral branch narrow, tubular with rounded apex, as long as sternal process; in ventral view sagittate, pointing anteriorly. Phallus, in lateral view shorter than segment IX, complex, ventral margin straight; in ventral view tubular, anterior end slightly wider than remainder.
Additional information. This species was referred to as "sp. 21" in Espeland and Johanson (2010 Diagnosis. Agmina circulata sp. nov. is distinguished from the other Agmina species in the genitalia by the presence of a pair of inferior appendages having a ventral part that, in lateral view, almost reaches as far posteriorly as the sternal processes, and with the apex of the sternal processes and inferior appendages that are separated by a rectangular incision. It resembles A. sagittata sp. nov. in the overall similarity in the genitalia, but A. circulata sp. nov. is smaller than A. sagittata sp. nov., and in the genitalia A. sagittata sp. nov. has superior appendages that are more rounded in lateral view, shorter sternal processes in relation to the inferior appendages, and the sternal processes of A. sagittata sp. nov. is slightly wider and straighter than those of A. circulata sp. nov. Etymology. Circulata from Latin, meaning rounded. Named for the parameres forming a circular loop at the apex. Measurements. Fore wing length 4.6 mm (N = 1). Total length of genitalia: 0.5 mm.
Description. Genitalia: In lateral view, segment IX narrowly bell-shaped anteriorly, apex located medially; in ventral view anteriorly concave lens-shaped with slightly rounded lateral margins. Sternal processes, lateral view, with each apex exceeding the length of tergum X and almost superior appendage, elongated narrowly triangular with acute apex, dorsal margin relatively straight, ventral margin gently concave; in ventral view, longer than inferior appendage, narrowing along their length, apex semi-acute, outer margin slightly rounded, inner margin straight. Tergum X in lateral view semi- triangular with concave anterior margin, dorsal margin with central bump, longer than superior appendage, longer than wide; in dorsal view, mesally separate, golf club shaped, inner margins forming shallow triangle. Parameres starting before tergum X, in lateral view long, slowly widening along 2/3 of its length, then narrow, apex forming ring, ending before apex of superior appendage; in dorsal view, separated, long, slender, straight, apex forming ring. Superior appendages, in lateral view, shorter than tergum X, irregularly rounded; in dorsal view stout, slightly curving inwards, with rounded outer margin and gently lobed inner margin, apex directed mesad with claw like process directed posteromesad. Inferior appendages, in lateral view, with posterad orientated short, rounded dorsal branch; ventral branch spine like, gently curving dorsad, almost reaching length of sternal process; in ventral view narrowly rounded base, slightly diverging along its length towards triangular apex. Phallus, in lateral view shorter than segment IX, tubular; in ventral view tubular with rounded anterior end.
Additional information. This species was referred to as "sp. 19" in Espeland and Johanson (2010 Diagnosis. Agmina digitata sp. nov. is distinguished from the other Agmina species by the combination of a pair of long, straight sternal processes that form almost a right angle with the anterior margin of segment IX in lateral view; the apex almost reaching the posterior terminal part of the superior appendages; inferior appendages well hidden behind the sternal processes in lateral view; and the superior appendages in lateral view being rhomboid and slightly produced dorsally at the posterior end. It resembles A. cunicula sp. nov. in the shape of the genitalia in lateral view, but A. digitata sp. nov. can be separated from A. cunicula sp. nov. by the shape and larger size of the superior appendages.
Etymology. Digitata, from Latin, meaning digit-like. Named for the shape of the apex of the superior appendage in dorsal view, which is toe-shaped.
Description. Genitalia: In lateral view, segment IX triangular anteriorly, apex located medially; in ventral view, anteriorly widely U-shaped. Sternal processes, lateral view, with each apex exceeding the length of tergum X and almost superior appendage, long, slender, dorsal margin with widely triangular notch before mid-length, then straight, ventral margin slightly concave, apex acute; in ventral view, almost twice as long as inferior appendage, long, slender, straight, slightly widening at club shaped, acute apex. Tergum X in lateral view approx. same length as superior appendage, irregularly triangular; in dorsal view, mesally separate, irregularly quadrilateral, rounded antero-mesally, inner margins forming U. Parameres starting before tergum X, in lateral view slender, with two thin whip-like processes at apex almost forming a ring, barely reaching superior appendage; in dorsal view, anteriorly separated, straight, posterior half fused to complex structure, apex with central lobe and truncate lateral branches ending at base of superior appendage. Superior appendages, in lateral view, axe-blade shaped with dorsal margin slightly concave, as long as tergum X; in dorsal view stout, curving inwards, with rounded outer margin and irregular inner margin, apex fingerlike, directed mesoposterad. Inferior appendages, in lateral view, with posterad orientated, small, widely triangular dorsal branch; ventral branch narrowly triangular, apex acute; in ventral view rounded at base, margins almost parallel, posterior margin concave with spine-like central process. Phallus, in lateral view shorter than segment IX, tubular; in ventral view tubular, anterior end with rounded lateral lobes, posterior end convex with spine-like lateral processes.
Additional information. This species was referred to as "sp. 50" in Espeland and Johanson (2010).  are orientated laterally before curving slightly mesally. It resembles many other species in the genus having small and almost rectangular or rhomboid superior appendages in lateral view, but is distinguished from these species by the presence of the lower parameres, as well as the straight sternal processes that are undulating in thickness. It resembles A. longicordata sp. nov. in the presence of a group of densely arranged setae at the posterior part of the parameres. Agmina longispina sp. nov. is easily separated from A. longicordata sp. nov. by the more rectangular shape of the inferior appendage plate in ventral view.
Etymology. Longispina, from Latin, longi and spina (noun, feminine) meaning long spine. Named for the shape of the sternal processes in lateral view, which are long and pointed posteriorly.
Description. Genitalia: In lateral view, segment IX triangular anteriorly; apex located dorsally; in ventral view anteriorly widely bell shaped. Sternal processes, lateral view, with each apex exceeding the length of apex X, almost reaching apex of superior appendage; wide at base, then long, slender, slightly curving ventrad, apex acute; in ventral view, longer than inferior appendage, slender, slightly converging towards acute apex. Tergum X in lateral view trapezoid, tapering along its length, slightly longer than superior appendage, longer than wide; in dorsal view, mesally separate, trapezoid, tapering along its length, inner margins forming bell. Parameres comprising pair of upper and lower branches; upper branches starting before tergum X, reaching base of superior appendage, in lateral view, first half very slender, then slightly broadening along its length, spine-like at apex with multiple wide setae on ventral surface, directed ventrad; in dorsal view, first half separated, slender, gradually converging and fusing at apex forming two large lobes; lower branches starting anteriorly of upper branches, slender and undulating in lateral view, in ventral view with anterior halves orientated posteriorly before bending laterally and slightly recurving mesally before posterior end, posteriorly exceeding superior appendages. Superior appendages, in lateral view, almost rectangular, slightly wider posteriorly, shorter than tergum Xin dorsal view stout, curving inwards, outer margin rounded, inner margin relatively rounded, apex clawlike, directed mesoanteriorly. Inferior appendages, in lateral view, with posterad orientated, slender, acute dorsal branch, exceeding length of superior appendage, ventral branch half the length of dorsal branch, broader, irregularly tapering along its length, abruptly narrowing at semi-acute apex; in ventral view widely bell-shaped at base, margins almost parallel, posterior margin straight with narrowly triangular central process. Phallus, in lateral view shorter than segment IX, slender, tubular, slightly curving downwards; in ventral view largely tubular, straight, slightly wider anteriorly.
Additional information. This species was referred to as "sp. 34" in Espeland and Johanson (2010). Diagnosis. Agmina magnahamata sp. nov. resembles many other Agmina species having short superior appendages with a large, strong mesally hook-shaped process that are present in other species, particularly in the Agmina sagittata species group. Agmina magnahamata sp. nov. resembles species with straight and posteriorly wide sternal processes, like A. digitata sp. nov. and A. circulata sp. nov., from which A. magnahamata sp. nov. is distinguished by the more complex parameres in the genitalia.
Etymology. Magnahamata, from Latin, magna and hamata (noun feminine), meaning large hook. Named for the process originating from the superior appendage, which in dorsal view is large, broad and hook-shaped.
Description. Genitalia: In lateral view, segment IX triangular anteriorly, apex located medially; in ventral view anteriorly widely U-shaped. Sternal processes, lateral view, with each apex exceeding the length of tergum X, almost reaching posterior margin of superior appendage; wide at base, then long, relatively slender, but slightly widening towards acute apex; in ventral view, almost twice as long as inferior appendage, parallel, slightly widening towards acute apex. Tergum X in lateral view largely rectangular, twice the length of superior appendage, longer than wide; in dorsal view, mesally fused, forming rounded lobe. Parameres starting before tergum X, reaching base of superior appendage, in lateral view, slender, slightly curving downwards, apex directed postero-dorsad, ventral surface with multiple wide setae directed ventrad; in dorsal view, first half parallel, long, slender, second half widening along its length, apex club shaped, on ventral surface multiple wide setae directed mesad. Superior appendages, in lateral view, largely quadratic, with large, broad hook-like process originating dorsally on posterior margin, curving downwards and back under main part, shorter than tergum X; in dorsal view stout, directed posterad, triangular, widening posteriorly, at posterior margin large hook-shaped protrusion with apex directed meso-anterad. Inferior appendages, in lateral view, with posterad orientated dorsal branch slightly longer than ventral branch, apices of both branches slightly converging, short, semiacute, dorsal margin convex, ventral margin slightly concave; in ventral view widely bell-shaped at base, margins parallel, posterior margin concave with triangular central process. Phallus, in lateral view shorter than segment IX, slender, tubular, slightly curving downwards; in ventral view tubular, relatively wide, lateral lobes at anterior end.
Additional information. This species was referred to as "sp. 42" in Espeland and Johanson (2010). Diagnosis. Agmina longicordata sp. nov. is distinguished from the other Agmina species by the inferior appendages in ventral view, which plate is narrowly heart-shaped and with a median process originating from the middle of its posterior margin. It is also unique by the presence of a group of small ventrally orientated setae densely arranged in a small circular area on each paramere. In addition, the sternal processes are almost boomerang-shaped, with each of the posterior part orientated in parallel. The shape of the plate formed by the inferior appendages is somewhat similar to that of A. sagittata sp. nov. and A. circulata sp. nov. but is easily separated from those species on the other characters. It resembles A. longispina sp. nov. in the presence of a group of densely arranged setae at the posterior part of the parameres. Agmina longicordata sp. nov. is easily separated from A. longispina sp. nov. by the oval shape of the inferior appendage plate in ventral view.
Etymology. Longicordata, from Latin, longi and cordata, meaning long and heart-shaped. Named for the shape of the inferior appendage, which is shaped like an elongated heart.
Material examined. Etymology. Cornuta, from Latin, meaning bill of a bird. Named for the shape of the inferior appendage in lateral view, which is long, beak-like.
Description. Genitalia: In lateral view, segment IX sharply triangular anteriorly, apex located medially; in ventral view anteriorly widely oblong. Sternal processes, lateral view, with each apex exceeding the length of superior appendage, narrowing along its length, curving downwards, apex acute, spine-like; in ventral view, robust, first half slightly diverging, second half converging, wider than first half, narrower blunt apices parallel with very straight inner margin, small spine-like process at apex, directed posteromesad. Tergum X in lateral view irregularly quadrilateral with concave anterior margin, tapering towards posterior, slightly longer than superior appendage; in dorsal view, mesally fused, inner margins forming wide U-shape with small, spinelike central process. Parameres starting around the base of tergum X, in lateral view, straight, tubular, with apex widening, sharply curving ventrad, ventral surface with large amounts of wide setae directed ventroanterad; in dorsal view, robust, finger-like, outer margins convex, inner margins concave, apex rounded with large amounts of mesally directed, wide setae on inner margin. Superior appendages, in lateral view, largely rectangular, longer than wide, slightly shorter than tergum X; in dorsal view almost parallel, outer margins straight, inner margins sigmoid; bent sharply mesally at apex, ending with claw-like structure directed mesoanterad. Inferior appendages, in lateral view, with posterad orientated, rounded, lobe-like dorsal branch, ventral branch longer, tapering along its length, ventral margin straight, convex towards acute apex; in ventral view elongated cordate, posterior margin concave with large triangular central process; Phallus, in lateral view approx. half the length of segment IX, dorsal margin sigmoid, ventral margin straight; in ventral view widest at mid-length, anteriorly with lateral lobes.
Additional information. This species was referred to as "sp. 29" in Espeland and Johanson (2010). Diagnosis. This species is very similar to A. semicampanula sp. nov., from which it is distinguished by the rectangular-shaped anterior margin of segment IX in ventral view; the angled sternal process in lateral view; and, in dorsal view, the pair of parameres that have sharply angled instead of rounded posteromesal corners.
Etymology. Campanula, named for the bell-shaped flowers in the genus Campanula, based on the shape of the inferior appendages in ventral view.
Description. Genitalia: In lateral view, segment IX sharply triangular anteriorly, apex located medially; in ventral view anteriorly obround. Sternal processes, lateral view, with each apex almost reaching apex of superior appendage, narrowing along its length, curving downwards, apex acute; in ventral view, robust, diverging, first 2/3 of equal width, apices parallel, narrowing, acuminate. Tergum X in lateral view irregularly quadrilateral with concave anterior margin, tapering towards posterior, approx. same length as superior appendage; in dorsal view, mesally fused, forming rounded lobe, in- ner margins forming narrow bell. Parameres starting before tergum X, in lateral view long, slender, straight, acute apex slightly directed dorsad; in dorsal view, long, slender, widening at truncated apex, posterior margin straight. Superior appendages, in lateral view, oblong, with posterodorsal indentation, approx. same length as tergum X; in dorsal view stout, almost semi-circular with rounded outer margin, inner margin semistraight, at apex large claw-shaped structure directed anteromesad. Inferior appendages, in lateral view, with posterad orientated rectangular dorsal branch and directly adjacent slender ventral branch of equal length, whole structure semi-rectangular; in ventral view bell-flower shaped. Phallus, in lateral view slightly shorter than segment IX, posterior half tapering along its length; in ventral view tubular with small lateral lobes at anterior end.

Additional information.
This species was referred to as "sp. 24" in Espeland and Johanson (2010 Measurements. Fore wing length 4.1 mm (N = 1). Total length of genitalia: 0.5 mm.
Description. Genitalia: In lateral view, segment IX triangular anteriorly, apex located dorsally; in ventral view anteriorly widely U-shaped. Sternal process es, lateral view, with each apex almost reaching apex of superior appendage, narrowing along its length, slightly curving downwards, apex acute; in ventral view, robust, anterior half diverging, posterior half parallel, tapering along their length, apex acuminate. Tergum X in lateral view irregularly quadrilateral with concave anterior margin and convex posterior margin, tapering towards posterior, approx. same length as superior appendage; in dorsal view, mesally fused, forming slightly irregular lobes, inner margins forming acuminate triangle. Parameres starting before tergum X, in lateral view long, slender, straight, bifurcating at apex, with dorsal branch orientated sharply dorsad and ventral branch orientated sharply ventrad; in dorsal view, long, slender, straight, slightly converging along its length, slightly diverging and widening at blunt apex. Superior appendages, in lateral view, oblong with anterodorsal tooth, approx. same length as tergum X; in dorsal view stout, almost semi-circular with rounded outer margin, inner margin slightly concave, apex with short, stout, claw-shaped structure directed anteromesad. Inferior appendages, in lateral view, with both branches forming a single posterad orientated, rectangular structure with slightly dentated, truncated, posterior margin; in ventral view semi-bell-shaped, posterior margin slightly convex with shallow teeth. Phallus, in lateral view slightly shorter than segment IX, largely tubular, ventral margin straight; in ventral view tubular with small lateral lobes at anterior end.
Additional information. This species was referred to as "FP2 Agmina sp. 24" in Espeland and Johanson (2010). Diagnosis. Agmina cunicula sp. nov. is distinguished from other Agmina species in having genitalia that are relatively simple, particularly in lateral view where they are composed of two visible branches, the upper including a large tergum X and the superior appendage, and the lower including the very broad sternal process that hide the inferior appendages and slightly exceed the superior appendages posteriorly. It resembles A. digitata sp. nov. but in lateral view A. cunicula sp. nov. have superior appendages that are rounded posteriorly, while trapezoid in A. digitata sp. nov. The inferior appendages of A. cunicula sp. nov. are very short and deeply divided longitudinally, while those in A. digitata form a wide and long plate that is not divided.
Etymology. Cunicula, from Latin, meaning lengthened. Named for the very long sternal processes.
Description. Genitalia: In lateral view, segment IX narrowly U-shaped anteriorly, apex located medially; in ventral view anteriorly U-shaped. Sternal processes, lateral view, with each apex exceeding apex of superior appendage, narrowing along its length, straight, apex rounded, ventral margin slightly sigmoid, dorsal margin straight; in ventral view, long, almost parallel, slightly diverging towards rounded apex, lateral margin straight, medial margin convex. Tergum X in lateral view jar-shaped, with dorsal and ventral margins gently convex posteriorly; longer than superior appendage; in dorsal view, mesally fused, with lateral margins straight, mesal margin convex, narrowing towards posterior; inner margins forming bell-shaped structure with distended oval base. Parameres starting before tergum X, in lateral view slender, slightly curving downwards and widening at truncate apex; in dorsal view, long, slender, first half converging, second half diverging, widening towards apex, apex truncated, almost triangular with mesal process shaped like double-spine orientated posteromesad. Superior appendages, in lateral view, oblong, with slightly straighter anterior margin, shorter than tergum X; in dorsal view stout, slightly converging towards apex, outer margin almost straight, inner margin slightly concave, apex with thick hook-shaped structure and small spine directed anteromesad. Inferior appendages, in lateral view, very short, rounded structure with acute apex, at apex large, hook-shaped structure almost as long as main appendage, parallel with ventral margin of sternal process, then sharply bending, with apex pointing posterad; in ventral view with two posteriorly orientated branches, remainder hardly visible under sternal process. Phallus, in lateral view slightly shorter than seg- ment IX, slender, tubular, with broad spine-like lateral lobe at anterior end; in ventral view tubular with anterior end slightly widening, truncate, with small spine-like lateral processes directed anterad.
Additional information. This species was referred to as "sp. 11" in Espeland and Johanson (2010 ing process, which make them easily recognised from other Agmina species. Agmina cerritula sp. nov. is distinguished from A. monstrosa sp. nov. by having less strongly narrowing sternal processes in lateral view and that the apex is more truncated, and the superior appendages are broader posteriorly compared to those in A. monstrosa sp. nov. Agmina recurvata sp. nov. have similar inferior appendages in lateral view, but in this species the long processes are paired, not simple as in A. serricula sp. nov. Etymology. Cerritula, from Latin, meaning weird. Named for the weirdly shaped inferior appendage. Measurements. Fore wing length 4.4 mm (N = 1). Total length of genitalia: 0.8 mm.
Description. Genitalia: In lateral view, segment IX sharply triangular anteriorly, apex located ventrally; in ventral view anteriorly oval. Sternal processes, lateral view, with each apex exceeding apex of superior appendage, robust, largely of equal width throughout its length, margins slightly sigmoid, apex widely rounded, ventrally with short spine-like process directed posteroventrad; in ventral view, very long, tapering along their length, margins almost straight, apex blunt with short, spine-like process directed posteromesad. Tergum X in lateral view saddle-shaped, anteriorly triangular with straight dorsal margin, slightly longer than superior appendage; in dorsal view, mesally fused, almost rectangular, inner margins forming wide V-shape. Parameres starting before tergum X, in lateral view slightly widening along its length, with convex dorsal margin until mid-length, then narrowing before again widening, forming long loop at apex running parallel with ventral margin, and reaching ventral posterior corner of superior appendage; in dorsal view, separated, slender, first 1/3 slightly diverging, second 1/3 widening and diverging, third 1/3 initially parallel, then curving laterally and looping anterad. Superior appendages, in lateral view, axe-blade shaped with posterior margin slightly dentate, slightly shorter than tergum X; in dorsal view very stout, equally long as wide, almost rectangular, apex truncated, mesally with stout claw-like process directed anteromesad, followed by two small triangular processes of diminishing size. Inferior appendages, in lateral view, slender, longer than sternal process, initially directed posteroventrad, then greatly curving, forming a wide U-shape, with acuminate apex directed dorsad posterodorsally of apex of sternal process; in ventral view long, slender, tubular with straight margins, longer than sternal processes, apex acute. Phallus, in lateral view almost as long as segment IX, slender, tubular, straight, with long spine-like process near anterior end; in ventral view tubular, slightly wider at base with rounded lateral lobes and sheet-like structure.
Additional information. This species was referred to as "sp. 46" in Espeland and Johanson (2010).

Agmina monstrosa sp. nov. http://zoobank.org/9D9A3F19-BD9D-4AE9-B343-0CFC9980750C Figs 84-88
Diagnosis. Both Agmina monstrosa sp. nov. and A. cerritula sp. nov. have genitalia with inferior appendages that form a very long, slender and dorsally curving process, which make them easily recognised from other Agmina species. Agmina monstrosa sp. nov. is distinguished from A. cerritula sp. nov. by having clearly narrowing sternal processes in lateral view and that the apex is tapering into a long, ventrally orientated tip, and the superior appendages are narrowing posteriorly instead of broadening as in A. monstrosa sp. nov.
Description. Genitalia: In lateral view, segment IX sharply triangular anteriorly, apex located ventrally; in ventral view anteriorly oblong. Sternal processes, lateral view, with each apex greatly exceeding apex of superior appendage, robust, widest at midlength, then tapering along its length, with straight margins, acuminate apex strongly curving posteroventrad; in ventral view, very long, nearly parallel, tapering along their length, posterior third uniformly narrow, with acuminate apex directed mesad. Tergum X in lateral view saddle-shaped, anteriorly semi-triangular with almost straight dorsal margin, slightly longer than superior appendage; in dorsal view, mesally fused, almost rectangular, inner margins forming bell. Parameres starting before tergum X, in lateral view wide at base, then slender, apex complex with multiple branches, possibly forming a loop, reaching apex of superior appendage; in dorsal view, separated, initially converging, then twisting and diverging, slightly widening along their length, apex forming large loop covered by superior appendage. Superior appendages, in lateral view, small, cone-shaped with convex anteroventral margin, slightly shorter than tergum X; in dorsal view very stout, slightly longer than wide, apex rounded, mesally with stout claw-like process directed anteromesad, followed by small triangular process and slightly larger claw-like process. Inferior appendages, in lateral view, slender, longer than sternal process, initially directed posteroventrad, then widely curving, forming a very wide U-shape, with acuminate apex directed almost dorsad posterodorsally of apex of sternal process; in ventral view long, slender, tubular with almost straight margins, longer than sternal processes, apex acuminate. Phallus, in lateral view shorter than segment IX, slender, tubular, straight, with long spine-like process near anterior end; in ventral view tapering along its length, widening at apex, two large rounded lateral lobes at anterior end.
Additional information. This species was referred to as "sp. 7" in Espeland and Johanson (2010). Diagnosis. The inferior appendages of A. rectangulata sp. nov. is unique from most other species in the genus due to being almost rectangular in ventral view; and a wide, with a convex ventral margin and presence of a posterodorsal and posteroventral process when viewed from the lateral side. The inferior appendages of Agmina diriwi A. hexacantha Ward, 2003 resemble those of A. rectangulata sp. nov. in lateral view, except lacking the posterodorsal processes, and in ventral view the shape of the appendages are different, i.e., the plate is produced posteriorly in A. rectangulata while excised in A. diriwi and A. hexacantha. Agmina rectangulata is morphologically closest to A. chela sp. nov. Agmina chela sp. nov. is distinguished from A. rectangulata sp. nov. by the inferior appendages having slightly larger posterodorsal process in lateral view; in ventral view, a more truncate posterior margin of inferior appendages. Furthermore, the superior appendage is clearly longer than in A. rectangulata sp. nov. in lateral view.
Etymology. Rectangulata, from Latin, meaning rectangular. Named for the shape of the inferior appendages having an almost rectangular impression. Measurements. Fore wing length 4.0-4.3 mm (N = 3). Total length of genitalia: 0.5 mm.
Description. Genitalia: In lateral view, segment IX triangular anteriorly, apex located medially; in ventral view anteriorly U-shaped. Sternal processes, lateral view, with each apex reaching apex of superior appendage, gently curving with convex dorsal margin and concave ventral margin, apex blunt; in ventral view, parallel, ventral margin convex, mesal margin straight, apex semi-acute. Tergum X in lateral view saddleshaped, anteriorly narrowly triangular with acuminate apex, anterior margin concave, dorsal margin straight, approx. same length as superior appendage; in dorsal view, mesally fused, irregular quadrilateral, inner margin widely U-shaped. Parameres starting at base of tergum X, in lateral view wider at base, then slender, tubular, second half extremely wide sheet-like, curving laterally and ventrally, with apex below superior appendage; in dorsal view, separated, wider at base, then slender, tubular, wider, clubshaped apex with lateral spine. Superior appendages, in lateral view, nearly oblong with straighter anterior margin, similar in length to tergum X; in dorsal view converging, longer than wide, finger-shaped with rounded apex with claw-like spine directed anteromesad, two small triangular processes on mesal margin anteriorly of claw. Inferior appendages, in lateral view, with dorsoposterad orientated shorter, claw-shaped dorsal branch; ventral branch longer, strongly curving at base, then straight, orientated dorsad; in ventral view almost rectangular with oval anterior margin, posterior margin widely triangular with ventral process forming acuminate apex, dorsal processes barely exceeding posterior margin, apices rounded with triangular notch mesally, lateral margins slightly undulating. Phallus, in lateral view shorter than segment IX, dorsal margin straight, ventral margin concave; in ventral view tubular, slightly tapering towards apex, with triangular lateral lobes near anterior end.
Additional information. This species was referred to as "sp. 45" in Espeland and Johanson (2010).

Agmina chela sp. nov. http://zoobank.org/7547FEDE-D338-4EB3-8B5D-DE125EBE80CB Figs 94-98
Diagnosis. The inferior appendages of A. chela sp. nov. are distinguished from those of most other species in the genus due to being almost rectangular in ventral view; and a wide, with a convex ventral margin and presence of a posterodorsal and posteroventral process when viewed from the lateral side. The inferior appendages of Agmina diriwi A. hexacantha Ward, 2003 resemble those of A. chela sp. nov. in lateral view, except lacking the posterodorsal processes, and in ventral view the shape of the appendages is different, i.e., the plate is produced posteriorly in A. chela while excised in A. diriwi and A. hexacantha. The species that is closest morphologically is A. rectangulata sp. nov. Agmina chela sp. nov. is distinguished from A. rectangulata sp. nov. in the inferior appendages by the, in lateral view, slightly larger posterodorsal process; in ventral view, a more truncate posterior margin of inferior appendages; and the superior appendage is clearly longer than in A. rectangulata sp. nov., as seen in lateral view.
Etymology. From Greek chela (noun, feminine), meaning pincer-like claw. Named for the crab-claw shaped inferior appendage in lateral view.
Description. Genitalia: In lateral view, segment IX triangular anteriorly, apex located medially; in ventral view anteriorly U-shaped. Sternal processes, lateral view, with each apex almost reaching posterior apex of superior appendage, gently curving with convex dorsal margin and concave ventral margin, apex blunt; in ventral view, long, slender, almost parallel, blunt apex. Tergum X in lateral view anteriorly narrowly triangular with acuminate apex, anterior margin straight, dorsal margin slightly concave, shorter than superior appendage; in dorsal view, mesally fused, semi-rectangular, inner margins forming very wide U-shape. Parameres starting before tergum X, in lateral view widening along its length, apex club-shaped, not exceeding superior appendage; in dorsal view, separated, straight, club-shaped apices slightly converging, with complex, sheet-like structure curving ventrally, ending just below superior appendage, anterior end semi-triangular. Superior appendages, in lateral view, longer than wide, anterior margin straight, apex rounded, slightly tapering along its length; in dorsal view robust, longer than wide, apex rounded with claw-shaped spine directed mes- oanterad. Inferior appendages, in lateral view, with dorsoposterad orientated slightly shorter, but broader straight dorsal branch; ventral branch longer, curving at base, then straight, orientated dorsoposterad; in ventral view almost rectangular with oval anterior margin, posterior margin almost straight, narrow ventral process with blunt apex, dorsal processes clearly exceeding posterior margin, apices rounded with rounded notch mesally, lateral margins straight. Phallus, in lateral view shorter than segment IX, dorsal margin concave, ventral margin almost straight; in ventral view uniformly tapering along its length, with narrow lateral lobes at anterior end.
Additional information. This species was referred to as "sp. 55" in Espeland and Johanson (2010).
Diagnosis. Agmina piscaria sp. nov. is distinguished from other Agmina species in combination of having a subapical instead of apical position of the median hooks of the superior appendage, dorsal branch of inferior appendages that are hooked mesally at their apex, seen in dorsal view, the shape of the plate formed by the inferior appendages that is narrowly parallelogram-shaped with a long central lobe, and long, straight sternal process. Also A. amieuensis sp. nov. and A. christinae sp. nov. have a subapical position of the mesal hooks of the superior appendages, but A. amieuensis sp. nov. is separated from A. piscaria sp. nov. by the tapering superior appendage in lateral view, the short sternal processes, and the different shape of the inferior appendage. Agmina christinae sp. nov. has a group of megasetae on the dorsobasal part of the superior appendages that are absent in A. piscaria sp. nov.
Etymology. Piscaria, from Latin, meaning fish-like. Named for the fish-shaped phallus in ventral view. Measurements. Fore wing length 4.5 mm (N = 1). Total length of genitalia: 0.6 mm.
Description. Genitalia: In lateral view, segment IX triangular anteriorly, apex located medially; in ventral view anteriorly oblong. Sternal processes, lateral view, with each apex almost reaching posterior apex of superior appendage, anterior half triangular, abruptly narrowing around mid-length, posterior half slender, straight with semi-acute apex, directed posteroventrad; in ventral view, anterior half robust, slightly diverging, posterior half narrower, almost parallel, apices blunt. Tergum X in lateral view anterodorsally narrowly triangular, with anterior margin concave, dorsal margin straight, ventral margin convex, shorter than superior appendage; in dorsal view, mesally fused, strongly concave anteriorly, inner margin forming U-shape. Parameres in lateral view anteriorly triangular, then abruptly narrowing, midpart very slender, curving, apex much wider, club-shaped, directed dorsad, with two warts at posterior margin; in dorsal view, widely separated, bifurcated anteriorly, slender, anterior half slightly diverging, posterior half parallel, narrowly, club-shaped at apex. Superior appendages, in lateral view, largely oval, longer than wide, with wide setae at posterior part of dorsal margin; in dorsal view robust, slightly converging, lateral margin straight, mesal margin convex with claw-like process directed anteromesad near rounded. Inferior appendages, in lateral view, with dorsoposterad orientated dorsal branch with rounded apex slightly longer than ventroposterad-directed narrow, acuminate, ventral branch; in ventral view rhomboid, with ventral branch forming long, acute, central process, club-shaped dorsal processes exceeding posterior margin. Phallus, in lateral view as long as segment IX, tubular, of equal width along its length, curving upwards towards posterior; in ventral view fish-shaped.
Description. Genitalia: In lateral view, segment IX narrowly U-shaped anteriorly, apex located mesally; in ventral view anteriorly ovoid. Sternal processes, lateral view, with each apex reaching posterior half of superior appendage, straight, directed posterad, narrowing along its length, apex acuminate. in ventral view, diverging, lateral margins convex, mesal margins slightly convex anteriorly, apices acuminate. Tergum X in lateral view triangular; in dorsal view, narrowly triangular with concave posterior margin. Parameres absent. Superior appendages, in lateral view, twice as long as tergum X, saddle-shaped with spine like ventral process, two tubular, curving mesal processes, the anterior largely parallel with dorsal margin, apex blunt, the posterior parallel with posterior margin with acute apex; in dorsal view long, anterior half straight, parallel, curved mesad at 90 degree angle at mid-length, tapering along its length, apex clawlike directed anterad; main branch bifurcating anteriorly, with slender mesal branch curving upwards with acuminate apex; second, narrowly triangular mesal branch originating between first mesal branch and the curve of main branch. Inferior appendages, in lateral view, with posterad orientated large, lobe-like, acute dorsal branch, ventral branch in the form of small tooth; in ventral view dorsal branch forming long, slender shape with convex margins and two slender, slightly converging posterior processes, ventral branch forming small central tooth. Phallus, in lateral view shorter than segment IX, tubular, straight, with anterior lateral lobes; in ventral view slightly wider posteriorly, thin lateral lobes anteriorly.
Additional information. This species was referred to as "sp. 8" in Espeland and Johanson (2010 Measurements. Fore wing length 3.2 mm (N = 1). Total length of genitalia: 0.6 mm.
Description. Genitalia: In lateral view, segment IX semi-triangular with rounded apex located medially; in ventral view anteriorly widely U-shaped. Sternal processes, lateral view, with each apex barely exceeding posterior apex of tergum X, ventral margin almost straight, dorsal margin with rectangular protrusion after mid-length, apex acuminate; in ventral view, parallel, triangular with acuminate apices, rounded lobes after mid-length on lateral margin. Tergum X in lateral view pot-shaped with anterior margin concave, posterior margin straight, higher than long; in dorsal view, irregularly quadrilateral with slightly concave posterior margin. Parameres long, in lateral view slender equally wide along their length, curving upwards towards narrowly clubshaped apex with megasetae at ventral margin, ending before apex of superior appendage; in dorsal view, anterior half slender, converging, posterior half diverging, apex complex sheet-like folding, with megasetae on mesal surface. Superior appendages, in lateral view, triangular, twice as long as tergum X, apex with short spine; in dorsal view stout, longer than wide, slightly widening along their length, abruptly narrowing towards posteromesad directed spine-like apex. Inferior appendages, in lateral view, with posterad orientated large, lobe-like, blunt dorsal branch, ventral branch in the form of small tooth; in ventral view dorsal branch forming slender shape narrowing along its length, with slightly convex margins, two slender, converging posterior processes, ventral branch forming central tooth. Phallus, in lateral view slightly shorter than segment IX, carafe-shaped, straight; in ventral view tubular, straight.
Additional information. This species was referred to as "sp. 15" in Espeland and Johanson (2010 A. cheirella Ward, 2003. The species in the nodosa species group are characterised by having relatively short inferior appendage that is simple and curving dorsally in lateral view.
Diagnosis. Agmina rostrata sp. nov. is characterised by having long posteriorly orientated sternal processes that are almost parallel-sided along their length, and heavily sclerotised parameres with posterior part orientated dorsally and deeply bifurcated in lateral view. Also A. dathioensis sp. nov., A. rougensis sp. nov. and A. lata sp. nov. have long and parallel-sided sternal processes in lateral view. Agmina rostrata sp. nov. is distinguished from A. dathioensis sp. nov. in that the sternal process is orientated posteriorly, not ventrally; and from A. lata sp. nov. in the slightly narrower sternal process in lateral view, and absence of megasetae on the posterior apex of the parameres on the mesal face of the superior appendages.
Etymology. Rostrata, from Latin, meaning beak-shaped. Named for the medial processes on the superior appendage in ventral view, together forming a beak-shaped process.
Description. Genitalia: In lateral view, segment IX sharply triangular anteriorly, apex located dorsally; in ventral view anteriorly shallowly oblong. Sternal processes, lateral view, with each apex almost reaching apex of superior appendage, gently tapering along its length, slightly anti-sigmoid, apex truncated; in ventral view, parallel, of similar width throughout its length, slightly bulging at mid-length, apex acute, directed mesad. Tergum X in lateral view trapezoid, shorter than superior appendage, in dorsal view, mesally fused, semi-rectangular with rounded posterior corners, inner margin forming shallow, wide U. Parameres in lateral view complex, multi-branched, anteriorly directed posteromesad, then turning posterodorsad at 90 degree angle, gradually widening with two-pronged apex directed dorsad, mesal branch originating close to 90 degree angle, with wide circular apex; in dorsal view, widest at mid-length, there with large open beak-shaped protrusion directed mesad. Superior appendages, in lateral view, irregularly axe-blade shaped with acute, tooth-shaped apex dorsally, dorsal mar- gin slightly concave, ventral margin convex; in dorsal view, obtuse triangular with apex directed mesally. Inferior appendages, in lateral view, with dorsoanterad orientated slender dorsal branch with hooked, acute apex, ventral branch directed dorsoposterad, wider, shorter, with semi-acute apex, both branches exceeding dorsal margin of sternal process; in ventral view slender, with convex lateral margins, narrow, rounded anteriorly, slightly widening towards mid-length, ventral branch forming two broad posterior processes with blunt apices. Phallus, in lateral view shorter than segment IX, slightly narrowing along its length, posterior end acute; in ventral view irregular.
Additional information. This species was referred to as "sp. 4" in Espeland and Johanson (2010).

Agmina dathioensis sp. nov.
http://zoobank.org/AB710193-1B39-4314-8501-7E1143986475 Figs 120-124 Diagnosis. Agmina dathioensis sp. nov. is characterised by having long posteriorly orientated sternal processes that are almost parallel-sided along their length. Also A. rostrata sp. nov., A. rougensis sp. nov., and A. lata sp. nov. have long and parallel-sided sternal processes in lateral view. Agmina dathioensis sp. nov. is distinguished from all these species by not having heavily sclerotised parameres. Agmina dathioensis sp. nov. is furthermore distinguished from A. rostrata sp. nov. and A. lata sp. nov. in that the sternal processes are orientated ventrally, not posteriorly, and from A. lata sp. nov. in the slightly narrower sternal processes in lateral view, and absence of megasetae on the posterior apex of the parameres on the mesal face of the superior appendages. Measurements. Fore wing length 3.9 mm (N = 1). Total length of genitalia: 0.5 mm.
Description. Genitalia: In lateral view, segment IX sharply triangular anteriorly, apex located medially; in ventral view anteriorly ovoid. Sternal processes, lateral view, with each apex almost reaching apex of superior appendage, slightly wider anteriorly, curving downwards, apex blunt forming three small lobes; in ventral view, long, robust, parallel, of equal with along their length, apex acute directed posteriorly. Tergum X in lateral view trapezoid, shorter than superior appendage, in dorsal view, mesally fused with wide bridge, wide rounded lobe, inner margin forming very shallow U. Parameres starting at tergum X, in lateral view complex folded structure with truncated apex exceeding dorsal margin of superior appendage; in dorsal view, slender, hardly discernible structure. Superior appendages, in lateral view, irregularly quadrilateral, apex, wide, truncated, almost twice the length of tergum X; in dorsal view relatively long, slender, slightly curving mesad, narrowing along its length, long acute apex sharply curving mesad. Inferior appendages, in lateral view, with dorsoanterad orientated dorsal branch, initially wider, then tapering towards apex, ventral branch with wide, clubshaped apex, both branches exceeding dorsal margin of sternal process, main structure exceeding ventral margin of superior appendage; in ventral view relatively short, with convex lateral margins, ventral branch forming two broad posterior processes with blunt apices. Phallus, in lateral view shorter than segment IX, almost straight; in ventral view irregular, with long, narrow lateral processes at posterior end.
Description. Genitalia: In lateral view, segment IX sharply triangular anteriorly, apex located dorsally; in ventral view anteriorly U-shaped. Sternal processes, lateral view, with each apex exceeding apex of superior appendage, anterior half wide, tapering along its length, posterior half, long, slender, spine like with acute apex; in ventral view, robust, twice as long as inferior appendage, tapering along their length, converging, apex acute. Tergum X in lateral view quadratic with mesoanterad corner drawn out, equally long as high, in dorsal view, mesally fused with wide bridge, wide rounded lobe with two short, tooth-like protrusion, inner margin forming very shallow, wide V. Parameres starting at tergum X, in lateral view, short structure approx. four times longer than wide, with truncated apex barely exceeding the dorsal margin of superior appendage; in dorsal view, slender, hardly discernible structure. Superior appendages, in lateral view, irregular posterior half wider than anterior half, apex with claw-like process dorsally, directed ventrad, more than twice the length of tergum X; in dorsal view relatively long, slender, slightly converging towards apex, narrowing along its length, short, acute, claw-like apex sharply curving mesad. Inferior appendages, in lateral view, with dorsoanterad orientated dorsal branch, initially wider, then tapering towards apex, ventral branch with wide, club-shaped apex, ventral branch exceeding dorsal margin of sternal process, main structure slightly exceeding ventral margin of superior appendage; in ventral view narrow anteriorly, widening until mid-length, then straight, slightly narrowing, with ventral branch forming two slightly narrowing processes with rounded apex and tooth orientated mesad. Phallus, in lateral view much shorter than segment X, tubular, almost straight; in ventral view tubular with anterior triangular lateral lobes.
Description. Genitalia: In lateral view, segment IX triangular anteriorly, apex located medially; in ventral view anteriorly U-shaped. Sternal processes, lateral view, with knife-shaped with anterior half slightly narrowing, posterior end slightly wider, apex rounded; in ventral view, robust, long, of equal width along their length, apex rounded, anterior half diverging, posterior half parallel. Tergum X mug-shaped, in lateral view longer than high; in dorsal view, mesally well-separated, much longer than wide, rounded apex. Parameres in lateral view long, slender, curving upwards, small dorsal lobe at mid-length, apex club-shaped with megasetae directed ventrad, then drawn out into long acuminate process orientated dorsad, barely reaching superior appendage; in dorsal view, separated, initially converging, second half diverging widely, apex club-shaped with megasetae directed mesad. Superior appendages, in lateral view, not exceeding the length of tergum X, irregularly quadrilateral with rounded apex, claw-like process mesally on posterior margin, directed anteromesad; in dorsal view short, stout, rectangular process with large megasetae orientated mesad at mid-length of inner margin, rounded apex with long, curving, spine-like process orientated mesad. Inferior appendages, in lateral view, forming a single rounded, club-shaped structure, gently curving upwards; in ventral view slender, with slightly undulating convex lateral margins, two slightly converging, posterior processes slightly narrowing along their length, with apex directed posterolaterad. Phallus, in lateral view much shorter than segment IX, tubular, straight; in ventral view slightly wider anteriorly, tubular.
Additional information. This species was referred to as "sp. 38" in Espeland and Johanson (2010 Measurements. Fore wing length 4.0 mm (N = 1). Total length of genitalia: 0.6 mm.
Description. Genitalia: In lateral view, segment IX rounded triangular anteriorly, apex located medially; in ventral view anteriorly U-shaped. Sternal processes, lateral view, with long, rectangular, of equal width throughout their length, dorsal margin widely triangular at mid-length, apex truncated; in ventral view, robust, slightly diverging, inner margin straight, outer margin widest before mid-length, apex truncated, slightly pointed mesally. Tergum X large, longer than wide, dorsal and ventral margins sigmoid, anterior margin concave; in dorsal view, mesally separate, longer than wide, posterior half rectangular, anterior part triangular. Parameres in lateral view long, slender, wider anteriorly, curving upwards toward apex forming looped structure not exceeding any margins of the superior appendage; in dorsal view, separated, anterior 2/3 parallel, diverging at posterior 1/3, then largely parallel, apex elongated club-shaped. Superior appendages, in lateral view, triangular, shorter than tergum X, mesal spine close to apex; in dorsal view short, triangular with convex lateral margin, apex rounded, long, thin, curved, spine-like process posteriorly on inner margin, directed mesad. Inferior appendages, in lateral view, forming a single knife-blade-shaped structure, with straight dorsal margin, convex ventral margin with shallow notch on anterior half; in ventral view slender, rounded, triangular at base, widest just before mid-length, then slightly tapering towards apex with two parallel posterior processes with rounded apex. Phallus, in lateral view much shorter than segment IX, tubular; in ventral view wider anteriorly.
Additional information. This species was referred to as "sp. 5" in Espeland and Johanson (2010). Diagnosis. This species is easily distinguished from other Agmina species in lateral view by the sickle shaped sternal processes, each being slightly curved ventrally and with a sharp apex. It also has a large, claw-like structure apically in the superior appendages and pointed megasetae at the apex of the parameres.
Etymology. From Latin falx (noun, feminine), meaning sickle. Named for the sickle-shaped sternal process in lateral view.
Description. Genitalia: In lateral view, segment IX rounded triangular with slightly upturned apex anteriorly, apex located ventrally; in ventral view anteriorly U- shaped. Sternal processes, lateral view, with sickle-shaped, curving downwards posteriorly, on dorsal margin rounded notch at mid-length; in ventral view, slender, parallel, spine-like with acute apex. Tergum X small, oblong with drawn out, ventroanterad corner, wider than long; in dorsal view, mesally fused, short, not much wider than mesal bridge, posterior margins slightly convex. Parameres originating before tergum X, in lateral view tubular, slightly wider anteriorly, then of equal width along its length, thin, tubular, blunt, process on ventral margin before mid-length; in dorsal view, anterior half tubular, nearly parallel, posterior part folding in on itself, apex truncated, mesally with long, straight, megasetae directed mesad. Superior appendages, in lateral view, curving with rounded, slightly dentate apex, dorsally with long, downwardscurving, spine-like process, exceeding main appendage; in dorsal view irregular, equally wide as high with almost straight posterolateral process, with dentate lateral margin, above this with posteromesad curving acute, spine-like process. Inferior appendages, in lateral view, forming single rounded, rhomboid process with rounded apex; in ventral view short, slender with convex margin, apex with two thin spine-like posterior processes with acute apex. Phallus, in lateral view much shorter than segment IX, tubular; in ventral view wider posteriorly, with looped lateral lopes anteriorly.
Description. Genitalia: In lateral view, segment IX almost rectangular, apex located dorsally; in ventral view anteriorly slightly undulating, almost straight. Sternal processes, lateral view, with straight, gently tapering along their length, apex semiacute; in ventral view, anterior half robust, diverging, of equal width throughout their length, abruptly narrowing slightly posteriorly of mid-length, posterior part clubshaped with rounded apex. Tergum X semi-rectangular with anterior margin concave and posterior margin convex; in dorsal view, mesally fused, forming rounded lobe with shallow, wide notch at apex. Parameres originating before tergum X, in lateral view long, slender, initially trifurcated, then slender, tubular directed posterodorsad, forming complex looped structure at apex; in dorsal view, separated, initially converging, then diverging, straight, slightly widening along their length, thin, whip-like apex curving anterolaterad. Superior appendages, in lateral view, rounded, rectangular with truncated apex, as long as tergum X; in dorsal view parallel, curving mesally towards apex, anterior end triangular, lateral margin convex, mesal margin straight, claw-like mesoanterad directed process at apex. Inferior appendages, in lateral view, forming single slightly narrowing process with rounded apex exceeding lateral margin of sternal process; in ventral view slender, anterior end widely triangular, lateral margins parallel anteriorly, slightly converging posteriorly, apex with two spine-like, thin posterior processes. Phallus, in lateral view as long as segment IX, tubular, straight; in ventral view wider anteriorly, tubular.
Description. Genitalia: In lateral view, segment IX widely bell-shaped, apex located medially; in ventral view anteriorly oval. Sternal processes, lateral view, with broad structure with semi-acute apex located ventrally, ventral margin slightly concave; in ventral view, slightly diverging, tapering along their length, apex rounded. Tergum X trapezoid with concave anterior margin; in dorsal view, mesally separate, longer than wide, inner margin convex, posterior margin slightly concave. Parameres originating at tergum X, in lateral view initially slender, widening to a folded, twisting, sheet-like structure at apex; in dorsal view, slender, gently widening along their length, initially diverging, posterior half converging, apex rounded, club-shaped. Superior appendages, in lateral view, triangular with dorsal margin convex, apex acute directed posteromesad, spine-like mesal process curving downwards with apex directed anteromesad; in dorsal view longer than wide, outer margin sigmoid, inner margin convex, two apices directed postero mesad, mesal process directed anteromesad posteriorly of both apices. Inferior appendages, in lateral view, with bifurcated anterior half, ventral process running parallel with, and exceeding ventral margin of sternal process, posterior part forming a single narrowing lobe with rounded apex exceeding posterior margin of sternal process; in ventral view posterior half forming lobe-like structure narrowing towards rounded apex. Phallus, in lateral view almost as long as segment IX, tubular, straight; in ventral view wider anteriorly, tubular.
Additional information. This species was referred to as "sp. 18" in Espeland and Johanson (2010 Measurements. Fore wing length 4.1 mm (N = 1). Total length of genitalia: 0.5 mm.
Description. Genitalia: In lateral view, segment IX widely bell-shaped, apex located medially; in ventral view anteriorly oval. Sternal processes, lateral view, with tapering along their length, ventral margin almost straight, apex slender, rounded; in ventral view, slightly diverging, of nearly equal width throughout their length, abruptly narrowing towards rounded apex. Tergum X almost quadratic with concave anterior and posterior margin, drawn-out anteroventral corner; in dorsal view, widely mesally separated, twice as long as wide, outer margins convex, inner margins straight, but slightly undulating. Parameres originating at tergum X, in lateral view forming complex structure with looped apex; in dorsal view, slender, anterior part slightly converg-ing, posterior part diverging, apex not readily visible. Superior appendages, in lateral view, exceeding length of tergum X, wider at base, then slender, spine-like, straight, with acute apex; in dorsal view long, slender, slightly curving, converging towards acute apex. Inferior appendages, in lateral view, with bifurcated anterior half, ventral process straight, exceeding ventral margin of sternal process, posterior part forming a single narrowing lobe with rounded apex directed posteromesad, exceeding posterior margin of sternal process; in ventral view slender with convex margin, two acute, thin, posterior processes at apex. Phallus, in lateral view not exceeding length of segment IX, tubular, slightly curving; in ventral view wider anteriorly, with irregular lateral processes at mid-length.
Additional information. This species was referred to as "sp. 40" in Espeland and Johanson (2010).

Agmina complexa sp. nov. http://zoobank.org/C8FC9FA5-E342-4FA7-9C98-F45945CE312A Figs 161-165
Diagnosis. In ventral view, the inferior appendages of Agmina complexa sp. nov. form a narrow plate with a posteroapical incision laterally flanged by a row of small setae, similar to those of A. spina sp. nov., A. guttata sp. nov., and A. amieuensis sp. nov. Agmina complexa sp. nov. are easily separated from the above similar species by the much longer and almost oval sternal process in lateral view, and the superior appendages are strongly modified and form a pair of very large, anteriorly curving hooks. Agmina amplexa sp. nov. also have strongly modified superior appendages but A. amplexa sp. nov. has three pairs of more or less mesally orientated hooks instead of a single pair of hooks that are directed anterad in A. complexa sp. nov., as seen in dorsal view.
Etymology. Named for the complex shape of the superior appendage. Measurements. Fore wing length 3.5-4.3 mm (N = 2). Total length of genitalia: 0.8 mm.
Description. Genitalia: In lateral view, segment IX narrowly triangular, apex located medially; in ventral view anteriorly elongated, U-shaped. Sternal processes, lateral view, with club-shaped, dorsal margin convex, ventral margin straight, apex rounded; in ventral view, robust, slightly diverging, equally wide along their length, apex truncated. Tergum X irregular quadrilateral with concave anterior margin, convex posterior margin, lateral margins almost straight; as wide as high; in dorsal view, mesally fused, triangular with acute apex. Parameres slender, starting before tergum X, in lateral view sigmoid, widening at apex with concave posterior margin; in dorsal view, separated, long, slender, straight, widening along their length, posterior half slightly wider anteriorly, then equally wide along their length, posteriorly abruptly curving lateroanterad, apex hook-shaped directed anterad. Superior appendages, in lateral view, initially triangular, greatly narrowing forming ridged, greatly curved, narrow process with acute apex directed mesad; in dorsal view forming large hook-like structures, with apices greatly curving, directed anteromesad, crossing mesally. Inferior appendages, in lateral view, with posterad orientated single branch, straight with truncate apex with dorsad directed tooth dorsally, ventral margin and apex exceeding margin of sternal process; in ventral view very slender, lateral margins convex, acute, narrow posterior processes at apex. Phallus, in lateral view tubular, straight; in ventral view slightly wider anteriorly, with triangular lateral lobes, tubular.
Additional information. This species was referred to as "sp. 22" in Espeland and Johanson (2010).

Species group 5, dognyensis-group
Included species in this group are: Agmina dognyensis sp. nov., A. mana sp. nov., A. anterohamata sp. nov., A. joycei , and A. curvatacua sp. nov. This is a strongly supported monophyletic group based on DNA data (Espeland and Johanson 2010a), but no morphological characters are unique for the group. Diagnosis. Agmina dognyensis sp. nov. is unique in the combination of having a long, narrow, almost straight superior appendage in lateral view together with a narrow and uniformly tapering sternal processes having a dorsad bend at mid-length. Being armed with a series of megasetae on the posterior part of both parameres is also unique for this species.
Description. Genitalia: Total length 1.2 mm. In lateral view, segment IX low, long, rounded anteriorly, apex located slightly above mid-height of genitalia; in ventral view, anterior incision widely and deeply U-shaped. Sternal processes, lateral view, with very long, basally wide, uniformly narrowing and slightly Z-shaped, apex long, smooth, pointed posterodorsally; in ventral view, undulating and posteriorly orientated along their length, parallel-sided except narrowing at apex. Tergum X long, narrowly triangular, with convex posterior and concave dorsal margin with dorsal process at mid-length; in dorsal view, mesally separate, mesal margin convex. Parameres weakly developed; anterior part slender in lateral view, starting well below and before tergum X, short anterior part orientated posteriorly before slightly curving dorsally below mid-length of tergum X; dividing into short dorsal branch fused with dorsobasal part of superior appendage, and robust ventral branch exceeding ventral part of superior appendage and with long row of long, posteriorly orientated megasetae; in dorsal view, separated and S-shaped at base, almost invisible along tergum X and superior appendage, except ventral branch and its mesally orientated megasetae. Superior appendages, in lateral view, simple, slender, with parallel, almost straight dorsal and ventral margins; apex with small right-angled incision; in dorsal view narrow, weakly diverging from mid-length; apex rounded. Inferior appendages, in lateral view, with wide, posterodorsal corner produced into short, narrow process; ends near basis of superior appendages; in ventral view short, almost oval shield with both posterodorsal and posteroventral corners separated mesally by U-shaped ventral and V-shaped incision. Phallus, in lateral view straight, approx. as long as tergum X, posteroventrally slightly produced posteriorly; in ventral view basis wide, gradually narrowing towards rounded apex.

Additional information.
This species was referred to as "sp. 37" in Espeland and Johanson (2010 Measurements. Fore wing length 5.1 mm (N = 1). Total length of genitalia: 0.9 mm.
Description. Genitalia: Total length 0.9 mm. In lateral view, segment IX low, long, rounded anteriorly, apex located dorsally in genitalia; in ventral view anterior incision narrowly deeply rectangularly shaped. Sternal processes, lateral view, with basally wide, sharply narrowing into dorsally curving, slender process, apex pointed; in ventral view, parallel-sided along their length before curving slightly laterally at four-fifth their lengths, apex pointing mesally. Tergum X small, triangular, with straight posterior and dorsal margin; in dorsal view, mesally separate, mesal margin not produces mesally. Parameres weakly developed; anterior part slender in lateral view, starting well below and before tergum X, short anterior part orientated anteriorly before looping posteriorly and with basal half running along dorsal margin of superior appendages, bending dorsally at mid-length, distal half slightly undulating dorsally, not exceeding superior appendages posteriorly; in dorsal view, fused at base, before separated and orientated posteriorly; Superior appendages, in lateral view, long, shaped like an open human hand in mesal view; divided into dorsal, posteriorly orientated branch and ventrally orientated branch near basis and orientated downwards; dorsal branch with several very long, ventrally orientated megasetae; ventral branch with approx. two long megasetae orientated posteriorly; in dorsal view narrow, weakly undulating and tapering posteriorly, slightly diverging along their length; megasetae on dorsal branch confined to mesal surface; ventral branches orientated posteromesally, almost dilated and with long megasetae orientated posteriorly. Inferior appendages, in lateral view, with wide, truncated posteriorly; ends well before apex of sternal processes; in ventral view long, narrow basally, slightly widening from 1/3 but narrowing gradually towards rounded apex. Phallus, in lateral view slightly longer than superior appendage, almost straight; in ventral view almost equally wide along its length.

Additional information.
This species was referred to as "sp. 36" in Espeland and Johanson (2010). length. Being armed with numerous megasetae on both dorsal and ventral branches of the superior appendage is also unique for this species.
Etymology. Antero and hamata, from Latin, meaning anterior and hook-shaped. Referring to the forwardly curved hook at the posterior end of the superior appendages.
Description. Genitalia: Total length 0.9 mm. In lateral view, segment IX low, long, rounded anteriorly, apex located dorsally in genitalia; in ventral view anterior incision widely and deeply U-shaped. Sternal processes, lateral view, with slightly arrowhead shaped, narrowing from mid-length, posterior part not produced posteriorly; in ventral view, slightly narrowing till mid-length, parallel-sided from mid-length and slightly diverging along their length, each with bifurcated apex. Tergum X trapezoid with sharply pointing anteriorly and rounded pointing posteriorly; dorsal margin straight; in dorsal view, mesally separate, mesal margin indistinctly triangular. Parameres anterior part slender in lateral view, starting well below and before tergum X, short anterior part orientated posteriorly, central part orientated posterodorsally, distal part divided into branch orientated posteroventrally and dorsal branch orientated posteriorly, not exceeding superior appendages posteriorly; in dorsal view, separated along their length, bifurcated at basis, orientated posterad, few stout setae present on dorsomesal faces immediately after end of tergum X; few long megasetae present in row on plate-like mesal process; distal part completely fused with superior appendages. Superior appendages, in lateral view, long, with posterodorsal corner strongly produced into strong anteroventrally orientated hook; posteroventral corner slightly produced posterad; dorsal margin almost straight; in dorsal view slender and almost parallelsided except narrowing towards apex; apex curving mesally and anteriorly. Inferior appendages, in lateral view, with short, finger-like dorsal branch orientated posterodorsally, posteroventral corner right-angled, without branch; in ventral view forming a nearly oval shield almost completely hiding dorsal branches; dorsal branches separated by narrow, shallow incision. Phallus, in lateral view approx. as long as inferior appendages, straight, with pair of posteriorly orientated spines situated on posteroventral margin; in ventral view wide along its length.
Additional information. This species was referred to as "sp. 3" in Espeland and Johanson (2010).

Agmina curvatacua sp. nov. http://zoobank.org/9DA8F862-84DA-4F36-BE63-DDD97B1046AE Figs 180-184
Diagnosis. Agmina curvatacua sp. nov. is unique among the Agmina species in that the pair of parameres form long, needle-shaped processed exceeding the posterior margin of the superior appendages; the basis of the inside of the superior appendages have the parameres with a cluster of posteromesad megasetae; and in lateral view, the sternal process tapers posteriorly and is curved ventrally before apex.
Etymology. Curvata and acua, from Latin, curved and needle-shaped. Referring to the curved needle-shaped posterior part of the sternal processes.
Description. Genitalia: In lateral view, segment IX evenly triangular anteriorly, apex located little below midheight of genitalia; in ventral view anterior incision widely and deeply V-shaped. Sternal processes, lateral view, with almost rectangular with straight dorsal and ventral margins, posterior margin convex; in ventral view, slightly narrowing and diverging along their length, with rounded, posteriorly directed subapical region, apex narrow and curving mesally. Tergum X in lateral view rounded triangular posteriorly, dorsal margin with dorsally orientated process approx. at mid-length; longer than high; in dorsal view, mesally separate, mesal margin almost sigmoid. Parameres anterior part slender in lateral view, starting well below and before tergum X, anterior part orientated posteriorly, central part orientated posterodorsally, distal part orientated posteroventrally and exceeding superior appendages posteriorly; in dorsal view, separated along their length, broad at basis, orientated posterad, few stout setae present on dorsomesal faces immediately after end of tergum X; long megasetae present in row on small plate-like mesal process; distal part separated from superior appendages and almost tangential before curving laterally before mesally pointing end. Superior appendages, in lateral view, almost rectangular with straight dorsal and ventral margins, posterior margin convex; in dorsal view forming parallelogram-shaped lobes with almost straight lateral and undulating mesal margins. Inferior appendages, in lateral view, with short, thick dorsal and ventral branches, dorsal branch curving slightly upward into triangular apex, ventral orientated posteriorly into rounded apex; ventral branch almost reaching as far posteriorly as sternal process; in ventral view with ventral branches forming narrowly diamond-shaped plate, almost arrow-like with rounded anterior and pointed posterior ends; dorsal branches divided by narrow, V- shaped incision above ventral branches. Phallus, in lateral view straight, approx. as long as width of genitalia; in ventral view widest immediately before mid-length, distal part almost parallel-sided.
Additional information. This species was referred to as "sp. 2" in Espeland and Johanson (2010).

Species group 6, bimaculata-group
Included species in this group are: Agmina bimaculata Schefter, 2000, A. recurvata sp. nov., A. taoensis sp. nov., A. triangulata sp. nov. andA. acula Ward & The species in the Agmina bimaculata species group all have inferior appendages that are more or less strongly expanded posteriorly and curving dorsally in lateral view. Diagnosis. Agmina recurvata sp. nov. is unique among Agmina species in having left and right inferior appendages longitudinally separated along their length, each with a small mesally orientated process, and a very long posterior branch that is strongly curving postero-dorsally before meeting basis of the superior appendages. Agmina taoensis sp. nov. also has long inferior appendages that are curving dorsally, but these are fused into a plate, as seen in ventral view. Furthermore, it lacks the mesally orientated processes, and the posterior branch is much shorter and not so strongly curved dorsally. Agmina cerritula sp. nov. and A. monstrosa sp. nov. have similar inferior appendages in lateral view, but in these species the long processes are united into a single central process, not paired and lateral as in A. recurvata sp. nov.
Description. Genitalia: In lateral view, segment IX sharply triangular anteriorly, apex located dorsally; in ventral view anterior incision widely and deeply U-shaped. Sternal processes, lateral view, with dilated posteriorly, dorsal and ventral margins almost straight; in ventral view, not clearly visible as it is hidden below inferior appendages. Tergum X in lateral view almost trapezoid, with dorsally orientated pro- cess at anterior corner; approx. as long as high; in dorsal view, mesally located close to each other but separate, anteromesal and posteromesal margins almost straight. Parameres anterior part slender, starting below and before anterior end of tergum X, almost straight till mid-height of posterior margin of tergum X, bending slightly ventrally and running into mid-part of superior appendage; in dorsal view, separated along their length, orientated posterad, equally narrow at basal half, few stout setae present on dorsal faces before end of tergum X; mesal lobe with setae present near basal part of superior appendage, fusing with superior appendage with long row of short megasetae. Superior appendages, in lateral view, dilated posteriorly, dorsal and ventral margins almost straight; in dorsal view forming oval lobes with almost straight lateral and convex mesal margins. Inferior appendages, in lateral view, with basal part narrowing posteriorly into very long and almost scale-like dorsally looped distal part; scale-like distal part orientated dorsally and lack setae; in ventral view separate along their lengths, not forming central plate, narrowly oval and divergent, posterior slander part orientated posteromesally. Phallus, in lateral view weakly sigmoid, approx. as long as inferior appendages; in ventral view vase shaped, with basal part almost double as wide as posterior part.
Additional information. This species was referred to as "sp. 28" in Espeland and Johanson (2010 Measurements. Fore wing length 3.2 mm (N = 1). Total length of genitalia: 0.4 mm.
Description. Genitalia: In lateral view, segment IX rounded triangular, apex located at mid-height of genitalia; in ventral view anterior incision widely and shallowly rectangularly shaped. Sternal processes, lateral view, with half circular, with almost straight dorsal margin and strongly convex ventral margin; in ventral view, almost equally wide along their length, with rounded apex, straight and diverging along their length. Tergum X in lateral view almost rectangular, with dorsally orientated process at anterior corner; slightly shorter than high; in dorsal view, mesally separate, mesal margin almost straight. Parameres slender, starting below anterior end of tergum X, almost straight till mid-height of basis of superior appendage, bending slightly ventrally before curving into posteroventral margin; in dorsal view, separated along their length, orientated posterad, equally narrow along their length, mesal lobe with setae present before superior appendage, fusing with superior appendage without long row of short megasetae. Superior appendages, in lateral view, half circular, with almost straight dorsal margin and strongly convex ventral margin; in dorsal view slightly widening along their length, each with rounded apex. Inferior appendages, in lateral view, with basal part almost rectangular, expanded into a prominent dorsally curving process being pointed apically; in ventral view almost diamond-shaped with sharply triangular lateral corners; with deep and narrow posterior incision, posterior branches orientated posteriorly and slightly diverging. Phallus, in lateral view slightly curving posteriorly at basis before sub-straight posterior main part; in ventral view vase shaped, with basal part almost double as wide as posterior part.
Additional information. This species was referred to as "sp. 6" in Espeland and Johanson (2010).

Agmina triangulata sp. nov.
http://zoobank.org/3780CF2B-093B-  Diagnosis. Agmina triangulata sp. nov. resembles A. comata Ward, 2003 in the genitalia, particularly, in lateral view, the triangular shape of the superior appendages, and the presence of a cluster of mesally orientated setae on the median face of the superior appendages; the curving sternal processes and inferior appendages. Agmina triangulata sp. nov. is distinguished from A. comata in that the superior appendages are slightly more strongly produced posteriorly, the wider sternal process in lateral view, the much shorter dorsally curving branch of the inferior appendage in lateral view, and the more widely round shape of the inferior appendage in ventral view.
Etymology. Triangulata, from Latin, meaning triangular. Referring to the shape of the superior appendages in lateral view. Measurements. Fore wing length 3.9 mm (N = 1). Total length of genitalia: 0.6 mm.
Description. Genitalia: In lateral view, segment IX almost triangular, apex located at mid-height of genitalia; in ventral view widely oval. Sternal processes, lateral view, with sharply triangular, longer than high; in ventral view, slender, with almost uniformly concave inner margin, apex slightly curving mesally. Tergum X triangular in lateral view, slightly longer than high; in dorsal view, almost tangential mesally at narrow apices, posteriorly convex and anteriorly concave margins; Parameres slender, starting below anterior end of tergum X, almost straight till mid-height of basis of superior appendage, bending strongly ventrally before fading into posteroventral margin; in dorsal view, separated along their length, orientated posterad, slightly narrowing, with mesal lobe with small setae present before superior appendage, fusing with superior appendage and with long row of short megasetae. Superior appendages, in lateral view, sharply triangular, longer than high; in dorsal view basally narrow, expanding mesally at mid-length; apex pointing mesally; lateral margins nearly straight. Inferior appendages, in lateral view, with basal part almost rectangular, posteroventral margin parallel with posteroventral margin of sternal process, posterior corner expanded into finger-like, dorsally curved process not reaching as far posterior as apex of sternal process; in ventral view large, oval, with deep central posterior V-shaped incision; lateral processes orientated posterad and slightly mesally curving. Phallus, in lateral view sub-straight, slightly longer than superior appendage; in ventral view almost uniformly wide along its length.
Additional information. This species was referred to as "sp. 14" in Espeland and Johanson (2010).

Agmina bleuensis
Description. Genitalia: In lateral view, segment IX almost constituting of sternal process only; anterior apex located at mid-height of genitalia; in ventral view an- teriorly without incision. Sternal processes, lateral view, with small, slender, approx. twice as long as high, with slightly concave dorsal margin, ventral margin convex; in ventral view, slender, orientated posteriorly along their length; apex pointed. Tergum X sharply tapering dorsally, higher than long; in dorsal view, mesally separate, each with irregular mesal margin. Parameres weakly developed, with small, narrow, ventral branch before superior appendages, continues into dorsal part of superior appendage before looping downwards into ventral margin of superior appendage; in dorsal view, slender, curving mesally and ending at mid-length of superior appendages; each with wart-like structure at mid-length with few short setae. Superior appendages, in lateral view, small, slender, approx. twice as long as high, with slightly concave dorsal margin, ventral margin convex; in dorsal view parallel-sided and slightly curved mesally along their length, rounded apex with mesal tooth. Inferior appendages, in lateral view, with broad basis and two long posteriorly orientated branches; dorsal branch slightly curving upwards, ending near mid-length of superior appendage, apex needle shaped; ventral branch uniformly thick, curving slightly outwards along its length; in ventral view with shield-shaped basis; dorsal branches located laterally of ventral branches, almost straight and orientated posterad along their length; ventral branches broad and curving mesally along their length, apex on each side well separated mesally; apex widely triangular; setae present on most parts. Phallus, in lateral view almost as long as segment IX, curving posterad at 1/4; in ventral view vase-shaped, with wide rounded basis and narrow posterior two-thirds.
Additional information. This species was referred to as "sp. 49" in Espeland and Johanson (2010).

Agmina touhoensis sp. nov. http://zoobank.org/F8EBA7E0-7450-49C0-85B4-96EA74EA5B15 Figs 206-211
Diagnosis. Agmina touhoensis sp. nov. resembles most other species in the species group, particularly A. bleuensis sp. nov. and A. hamata due to the wide separation of the dorsal and ventral branches of the inferior appendages in lateral view. Agmina touhoensis sp. nov. is distinguished from A. bleuensis sp. nov. by the much longer sternal processes, and a dorsal branch of the inferior appendages that are almost as long as the ventral branches. It is distinguished from A. hamata by the wider superior appendages in lateral view, and the dorsal branch of inferior appendages that is almost straight instead of curving ventrally.
Description. Genitalia: In lateral view, segment IX almost constituting of sternal process only; anterior apex located immediately below mid-height of genitalia; in ventral view anteriorly without incision. In lateral view, sternal processes with small, almost straight dorsal margin, ventral margin pointing into triangular; in ventral view, slender, orientated posteriorly before strongly curving mesally before apex. Tergum X almost rectangular, with pointed anterodorsal corner; in dorsal view, mesally well separate, each forming narrow plate curving inwards, with slightly convex inner mar- gin. Parameres robust, tubular, starting below tergum X and ending at basal 1/3 of superior appendages, hidden behind upper branch of inferior appendages; in dorsal view, originating from transverse bow-like basis; orientated posteriorly, almost parallel, row of short megasetae present immediately after mid-length. Superior appendages, in lateral view, small, with almost straight dorsal margin, ventral margin pointing into triangular; in dorsal view bean-shaped, orientated posterad along their length, except slightly mesally orientated, rounded corners. Inferior appendages, in lateral view, with broad basis and two long posteriorly orientated branches; dorsal branch slightly undulating, ending near basis of superior appendages, apex needle shaped; ventral branch uniformly thick, curving outwards along its length; in ventral view with shield-shaped basis; dorsal branches located laterally of ventral branches, undulating and orientated posterad along their length; ventral branches broad and curving mesally along their length and almost tangential apically; apex narrowly triangular; setae confined to distal two-thirds. Phallus, in lateral view approx. half as long as segment IX, curving downwards at mid-length; in ventral view vase-shaped, with wide rounded basis and narrow posterior half.
Additional information. This species was referred to as "sp. 23" in Espeland and Johanson (2010). Diagnosis. The long and dorsally curving inferior appendages of A. wardi sp. nov. makes it resembling A. recurvata sp. nov., A. cerritula sp. nov. and A. monstrosa sp. nov. It is distinguished from the two latter by having paired processes of inferior appendages instead of a simple process; and from A. recurvata sp. nov. by the posteriorly narrowing sternal processes that end in a thick, straight apical megaseta that is orientated posteroventrally. Measurements. Fore wing length 3.8-4.0 mm (N = 3). Total length of genitalia: 0.6 mm.
Description. Genitalia: In lateral view, segment IX almost constituting of sternal process only; anterior apex located immediately below mid-height of genitalia; in ventral view anteriorly with widely U-shaped incision. Sternal processes, lateral view, with large, half circular with almost straight dorsal margin, without spines; in ventral view, slender and curving mesally from mid-length. Tergum X almost rectangular with posterodorsal rounded corner and anterodorsal pointed corner, in lateral view higher than long; in dorsal view, mesally situated closely but separate, each forming trapezoid lobes with straight inner margin. Parameres starting from lower anterior part of tergum X, narrow, bifurcating into dorsal and ventral branches basally in superior appendage; in dorsal view, forming pair of narrowing rays, each with small megasetae in short row located midway on mesal margin. Superior appendages, in lateral view, large, half circular with almost straight dorsal margin, without spines; in dorsal view uniformly wide along their length, parallelogram-shaped; apex strongly inwardly pointed. Infe- rior appendages, in lateral view, with broad basal part covered by setae separated from L-shaped mid-part that is smooth basally and with setae distally; apparently with a distal joint as long as height of tergum X; in ventral view with widely triangular basal part and truncate posterior corner; central part orientated laterally before looping mesally at mid-length; apex almost rounded. Phallus, in lateral view slightly shorter than length of segment IX, straight; in ventral view vase-shaped, with wide basis and narrow posterior half.
Additional information. This species was referred to as "sp. 39" in Espeland and Johanson (2010).
Agmina parallela sp. nov. http://zoobank.org/581632F3-2640-4D13-ACD9-41B8676FFE24 Figs 217-221 Diagnosis. Agmina parallela sp. nov. unique among Agmina species in the superior appendage that in lateral view is almost as large as segment IX and segment X combined, and is rounded club-shaped ventrally. In addition, the inferior appendages form a long ventral plate that is almost parallel-sided along its length.
Etymology. Parallela, derived from parallel, referring to the inferior appendages having almost parallel-sided lateral margins in ventral view.
Description. Genitalia: In lateral view, segment IX widely rounded anteriorly, almost trapezoid, apex located dorsally; in ventral view anteriorly with widely and shallow U-shaped incision. Sternal processes, lateral view, with very large, downwardly club-shaped, posterior and anterior margins almost parallel, apex widely rounded; in ventral view, absent. Tergum X deeply concave dorsally, posteriorly expanded dorsad into pointed triangular, in lateral view approx. as long as high; in dorsal view, forming small plates widely separated mesally. Parameres dorsally membranous, ventrally forming strongly sclerotised spines reaching to half-length of superior appendages; in lateral view slightly curving posteriorly; in dorsal view, separate and re-curved as basis, needle-shaped, almost straight after basis and pointing mesally. Superior appendages, in lateral view, very large, downwardly club-shaped, posterior and anterior margins almost parallel, apex widely rounded; in dorsal view narrow at basis, widening into mesal plates at mid-length, small mesally orientated tooth present at basis; row of long apical megasetae situated on inner surface and orientated mesally. Inferior appendages, in lateral view, with posterad orientated long dorsal branch with pointed apex; dorsal branch widely separated from ventral branch; ventral branch running parallel with dorsal branch, approx. double the width of dorsal branch and gently curving dorsally along its length; apex narrowly rounded; in ventral view rectangular plate-like ventral branch hiding dorsal branches, dorsal branches orientated posteriorly, each uniformly narrowing into acute apex. Phallus, in lateral view as long as segment IX, slender and slightly curving upwards; in ventral view equally wide along its length, double as wide as high.
Description. Genitalia: In lateral view, segment IX almost truncate anteriorly, apex located dorsally; in ventral view with anterior incision widely and deeply Vshaped. Sternal processes, lateral view, with large, oval, somewhat stalked basally, almost symmetrical dorsal and ventral sides; in ventral view, wide basally, narrowing to midway, almost parallel-sided along distal half; orientated posteriorly. Tergum X strongly concave before pointing dorsally, in lateral view slightly higher than long; in dorsal view, mesally separate, each forming short lobes with rounded margins. Parameres starting from anterior part of tergum X, posteriorly with small membranous dorsal part and sclerotised ventral part, ventral part with cluster of megasetae visible above base of superior appendages on mid-length of inner margin, a hook-like curved spine situated apically on each paramere and situated on the inside of superior appendages; in dorsal view, anterior parts fused into a V-shaped suture, basal megasetae almost meeting mesally, distal hooks curving inwards along their length, situated immediately anteriorly of large mesal processes. Superior appendages, in lateral view, large, oval, somewhat stalked basally, almost symmetrical dorsal and ventral sides; in dorsal view slightly widening, straight, and converging along their length, with long, mesal process before apex. Inferior appendages, in lateral view, with two posteriorly orientated almost parallel branches in lateral view, dorsal branch with a long and thick posteriorly orientated seta on ventral margin, apex acute; ventral branch as long as dorsal branch and with pointed apex; in ventral view, ventral branch with almost square basal half and single-rayed, narrow, tapering distal half; dorsal branches forming a pair of narrow triangular processes curving mesally at apex. Phallus, in lateral view slightly shorter than segment IX, slender and slightly curving upwards; in ventral view with triangular lateral processes on anterior half, distal half approx. equally wide.
Description. Genitalia: In lateral view, segment IX widely rounded anteriorly, apex located dorsally; in ventral view with anterior incision narrowly U-shaped, almost semi-oval. Sternal processes, lateral view, with shorter than high, slightly trapezoid; in ventral view, not clearly set off from rest of segment IX. Tergum X higher than long, convex dorsally and with small tapering dorsal process, in dorsal view, separate but almost tangential mesally, with rounded inner margins. Parameres robust, starting inside basis of tergum X, dividing before superior appendages into long, narrow and pointed dorsal branch and short, pointing ventral branch, both branches ending before midlength of superior appendage; in dorsal view, strongly fused and indistinguishable from superior appendage. Superior appendages, in lateral view, shorter than high, slightly trapezoid; in dorsal view almost parallelogram-shaped, with near parallel lateral and mesal margins; strong triangular mesally orientated tooth situated near apex. Inferior appendages, in lateral view, with almost parallel dorsal and ventral margins, posteriorly divided into short narrow triangular dorsal and almost equally short similarly shaped ventral branches widely separated by U-shaped cleft; in ventral view large, longer than wide, with convex lateral margins, narrowing at three-fourth its length; apex with central cleft; dorsal branches visible through the ventral branch, forming pair of wellseparated triangular processes. Phallus, in lateral view, slightly shorter than segment IX, uniformly slender along its length, straight; in ventral view with triangular lateral processes at basal 1/3; distal two-thirds with slightly concave lateral margins.
Description. Genitalia: In lateral view, segment IX narrowly rounded anteriorly, apex located dorsally; in ventral view anterior incision widely U-shaped. Sternal processes, lateral view, with large, almost half-spherical with straight dorsal margin; in ventral view, slender, basal part slightly orientated mesally, bent posteriorly at midlength. Tergum X sharply pointing dorsally, higher than long in lateral view; in dorsal view, mesally separate, each forming short lobes with almost straight inner margins. Parameres weakly developed, starting at basis of tergum X; in lateral view, bending upwards and meeting superior appendage basally at dorsal margin; in dorsal view, almost invisible, with group of stout, strictly mesally orientated megasetae. Superior appendages, in lateral view, large, almost half-spherical with straight dorsal margin; in dorsal view, narrowly parallel-sided along their length, smoothly convex lateral margins; apex with stout, finger-like spine pointing mesally. Inferior appendages, in lateral view, with ventral margin uniformly convex along its length; dorsal branch orientated strictly upwards, sharply triangular, with very long posterior margin ending in well separated ventral branch; ventral branch curving upwards along its length, with acute apex; in ventral view, with large central plate being approx. as long as wide, with convex lateral margins and triangular anteriorly; posteriorly with pair of long, widely separated processes curving mesally along their length. Phallus, in lateral view, much shorter than segment IX, straight; in ventral view, triangular with rounded anterior margin and pointed posteriorly.
Additional information. This species was referred to as "sp. 25" in Espeland and Johanson (2010 Measurements. Fore wing length 5.0 mm (N = 1). Total length of genitalia: 0.7 mm.
Description. Genitalia: In lateral view, segment IX narrowly rounded anteriorly, apex located at mid-height of genitalia; in ventral view, anterior incision widely Ushaped, with slightly mesally orientated anterior corners. Sternal processes, lateral view, with large, oval, slightly deeper convex ventrally than dorsally; mesal processes exceeds apex; in ventral view, basally wide, tapering and curving inwards along their length. Tergum X sharply pointing dorsally, higher than long in lateral view; in dorsal view, mesally separate, each forming short lobes with mesally pointing anterior corner. Parameres weakly developed, starting at lower basis of tergum X; in lateral view, bending upwards and with downwardly curving rounded apex; in dorsal view, almost invisible, with group of stout, antero-mesally orientated megasetae. Superior appendages, in lateral view, large, oval, slightly deeper convex ventrally than dorsally; mesal processes exceeds apex; in dorsal view, narrowly parallel-sided and curving inwards along their length, smoothly convex lateral margins; apex with stout, finger-like spine pointing mesally. Inferior appendages, in lateral view, with basal half of ventral margin almost straight, distal half uniformly convex; dorsal branch orientated strictly upwards, widely triangular, with concave posterior margin ending in well separated ventral branch; ventral branch wide at base, tapering and curving upwards along its length, with acute apex; in ventral view, with large central plate being approx. as long as wide, with angled lateral margins and triangular anteriorly; posteriorly with pair of long, widely separated processes orientated posteriorly along their length. Phallus, in lateral view approx. as long as segment IX, straight; in ventral view, triangular with rounded anterior margin and pointed posteriorly.
Additional information. This species was referred to as "sp. 9" in Espeland and Johanson (2010).

New species records
The following records are herewith recorded as new. The material is deposited at NHRS.

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In ventral view, plate of inferior appendages with lateral margins almost parallel-sided or diverging along its length (as in Figs 49, 71); in lateral view, sternal process not bifurcated (Fig. 69)  In lateral view, sternal process straight and thin; in ventral view, plate of inferior appendages with lateral margins almost parallel-sided along its length (as in Fig. 49 In ventral view, plate of inferior appendages with long lateral processes (as in Fig. 28 In lateral view, inferior appendages divided into a dorsal and a ventral branch (Fig. 206)  In lateral view, sternal process pointing posteroventrally (Fig. 195)  In ventral view, plate of inferior appendages generally oval with deep posterior incision (Fig. 198); superior appendages with dense area of small setae on mesal face (Fig. 196)  In lateral view, sternal processes almost parallel-sided along their length (Fig. 120)  In lateral view, sternal processes strongly narrowing at mid-length (Fig. 125)  In lateral view, superior appendages bifurcating (Fig. 105); sternal processes almost straight (Fig. 105) (Figs 115, 166); sternal processes slightly sigmoid (Fig. 115, 166)  Superior appendages each with mesad part approx. as long as posterad part (Fig. 106)  In lateral view, sternal processes broad, almost parallel-sided along their length, with truncate apex (Fig. 115)  In lateral view, superior appendages slender, almost parallel-sided along their length (Fig. 166); paraproct with row of posteroventrally orientated megasetae (Fig. 166)  In lateral view, superior appendages high, triangular (Fig. 115); paraproct without megasetae (Fig. 115)  ; superior appendages without long, slender ventrally orientated process (Fig. 41), but a hook-like spine can be present (Fig. 130)  In lateral view, sternal processes almost straight along their length (Fig. 41); inferior appendages hidden behind sternal processes (Fig. 41); in ventral view, plate of inferior appendages with narrow, spine-like central posterad process (Fig. 43)  In lateral view, sternal processes slightly bent ventrally at mid-length (Fig. 64); inferior appendages visible below sternal processes (Fig. 64); in ventral view, plate of inferior appendages with very wide central posterad process (Fig. 66)

27
Superior appendages apically with large, claw-like spine approx. as long as rest of superior appendages (Fig. 53)  In lateral view, tergum X longer than superior appendages (Figs 74, 135) ....30 29 Superior appendages with apical spine almost as long as rest of the superior appendages (Fig. 131); in ventral view, sternal processes almost parallel-sided along their length (Fig. 132)  Superior appendages with apical spine approx. 1/3 the length of the rest of the superior appendages (Fig. 147); in ventral view, sternal processes divided into an almost parallel-sided basal half and a oval distal half (Fig. 148)  In dorsal view, apical spines of superior appendages very thin, needle-like (Fig. 136); in lateral view inferior appendages surpassing posteriorly midlength of sternal processes (Fig. 135)  In dorsal view, apical spines of superior appendages very broad, hook-like (Fig. 75); in lateral view inferior appendages not surpassing posteriorly midlength of sternal processes (Fig. 74)  In lateral view, superior appendages with almost straight dorsal margin (Fig. 232)  In lateral view, sternal process sigmoid (Fig. 237)  In lateral view, sternal processes with triangular apex (Fig. 185)  In lateral view, sternal processes with finger-like apex (Fig. 185)  In lateral view, sternal process with finger-like apex shorter than maximum thickness of inferior appendages (Fig. 79)  -In lateral view, sternal process with finger-like apex more than 2 times longer than maximum thickness of inferior appendages (Fig. 84)  In ventral view, plate of inferior appendages with posterolateral branches along ventral margin separated by wide and deep incision (Figs 3, 7, 192 In ventral view, plate of inferior appendages with posterolateral branches along ventral margin tapering posteriorly (Fig. 192); mesal process of superior appendages orientated mesally (Fig. 191)  In lateral view, inferior appendages each with a dorsal and a ventral branch, and the dorsal branch is more than half the length of the ventral branch (Figs 201, 206)  In lateral view, each inferior appendage with dorsal branch longer than ventral branch (Fig. 15) Ward and Schefter (2000)); inferior appendages with ventral branch forming a single plate ( fig. 28 in Ward and Schefter (2000)) ... A. padi In lateral view, superior appendages narrowing posteriorly (Fig. 15); inferior appendages with ventral branch forming a pair of processes (Fig. 17) ..........48 48 In lateral view, parameres with a long process expanding below basis if superior appendages (Fig. 15)  In lateral view, each inferior appendage with dorsal and ventral branches narrowly separated at bases (figs 17 and 20 in Ward and Schefter (2000) In lateral view, sternal processes slightly curving ventrally along their length (Fig. 201); paraprocts without ventrally orientated megasetae under basis of superior appendages (Fig. 206)  In lateral view, sternal processes long, exceeding basis of superior appendages (Fig. 212)  In lateral view, superior appendages almost three times longer than high (Fig. 20); inferior appendages high along their length (Fig. 20); in ventral view, plate of inferior appendages with posterior incision shallower than half length of plate (Fig. 22)  In lateral view, superior appendages approx. two times longer than high (Fig. 25); inferior appendages high only at basis (Fig. 25); in ventral view, plate of inferior appendages with posterior incision deeper than half length of plate (Fig. 28)  In lateral view, sternal processes uniformly curving ventrally along their length (Fig. 50)  In lateral view, sternal processes bending in right angle and almost straight after bending point (Fig. 212)  In lateral view, superior appendage more than 3 times longer than high (Fig. 156)  In lateral view, superior appendage about 2 times longer than high (Ward 2003: fig. 5); in lateral view, inferior appendages with three finger-like posterior processes (Ward 2003: fig. 5 In lateral view, sternal processes with posterior 1/3 uniformly slender and strongly curving ventrally (Fig. 180); paraproct exceeding superior appendages posteriorly (Fig. 180)  In lateral view, superior appendage shorter than tergum X (Fig. 31)  In lateral view, superior appendage longer than tergum X (Figs 99, 175) ....66 66 In ventral view, plate of inferior appendages with long posterior central process (Fig. 102)  In ventral view, plate of inferior appendages without posterior central process (Fig. 177)  In lateral view, inferior appendages constituting a prominent dorsal and ventral branch (Fig. 222); superior appendages ovoid (Fig. 222); in ventral view, plate of inferior appendages with posterior central process longer than rest of the plate (Fig. 224)  In lateral view, superior appendages almost rectangular (Fig. 227); inferior appendages with dorsal and ventral branches narrowly separated (Fig. 227); in ventral view, plate of inferior appendages with convex lateral margins (Fig. 229)  In ventral view, plate of inferior appendages with undivided posterior central process (Fig. 102)  Superior appendages each with single, spine-like mesal process curving mesally along its length (Fig. 152); parameres without megasetae ....A. amieuensis sp. nov. -Superior appendages each with weakly defined spine-like mesal process straight and orientated mesoposteriorly along its length (Fig. 111); posterior end of parameres with cluster of megasetae (Figs 110, 111) ..........A. caraffa sp. nov. 73 In lateral view, superior appendages with dorsal branch wider than ventral branch (Fig. 105); inferior appendages situated dorsally of sternal processes (Fig. 105) (2003)); inferior appendages situated ventrally of sternal processes : fig. 56) In lateral view, superior appendages with hook-shaped apex (Fig. 175)

Discussion
The first Trichoptera from New Caledonia were described by Kimmins (1953), and including the species described here this number has now risen to 286, of which all but four species are endemic. The total diversity is expected to be even higher Johanson 2010a, Johanson and, and more work is needed to fully understand the biodiversity of the New Caledonian islands. Currently, very little is known about the early stages of New Caledonian species, and nothing at all is known about the early stages of members of the Agmina or Caledomina (Mary 2017). Like other members of the Ecnomidae, larvae are probably predatory and construct tubes of silk and fine sands, attached to various substrates. Associating adults and larvae using DNA barcoding (Johanson 2007) would be an important next step to increase the knowledge about Agmina and New Caledonian Trichoptera as a whole, and would make them available as valuable indicators for freshwater quality.