Corresponding author: Wenliang Li (
Academic editor: R. Meier
Four species of
Li W, Chen X, Yang D (2020) Four new species of the subgenus
The subgenus
It is controversial whether
The subgenus is recognized by anterior edge of frons glossy and upturned relative to rest of frons, face and parafacialia with gray pruinosity, shiny frons, hyaline yellow wings, scutum with intra-alar seta, katepisternum with a discal katepisternal seta, and mesonotum with 0–1 + 2–3 dorsocentral setae and 0–1 + 2–4 acrostichal setulae; most species have one to three pairs of strong acrostichal setulae (
General terminology follows
1 | Frons shining and face flat; mesonotum with 0–1 + 2–3 dorsocentral setae; katepisternum with one katepisternal seta; male genitalia: dorsal aedeagal sclerite present |
subgenus |
– | Frons often dull and face slightly concave; mesonotum with 0+3 dorsocentral setae; katepisternum with two katepisternal setae; male genitalia: dorsal aedeagal sclerite absent (if dorsal aedeagal sclerite present, but no presutural dorsocentral setae | 2 |
2 | Face with one pair of distinct oblong tumour processes on lower margin | 3 |
– | Face flat, without tumour process on lower margin | 4 |
3 | Wing black at base; arista long plumose, the longest ray longer than wide of first flagellomere; postgonite with four asymmetric long or short sclerites |
subgenus |
– | Wing yellow at base; arista short plumose, the longest ray as long as half width of first flagellomere; postgonite with two asymmetric long or short sclerites |
subgenus |
4 | Arista pubescent (some bare or short hair-like), wing yellow at base; postgonite consisting of one pair of sclerites, surrounding membranous aedeagus; aedeagal dorsal sclerite absent |
subgenus |
– | Arista short plumose, wing dark yellow or pale brown at base; postgonite consisting of one pair of narrow and asymmetric sclerites in ventral view; aedeagal dorsal sclerite broad, quadrate, rectangular, triangular or trapeziform |
subgenus |
5 | Mesonotum with 1+3 dorsocentral setae, with long acrostichal setulae | 6 |
– | Mesonotum lacking a presutural dorsocentral setae, without long acrostichal setulae | 7 |
6 | Arista bare; acrostichal setulae in two rows; male genitalia: anterior epandrium narrow and posterior broad, with a deep concavity on anterior ventral margin and a digitiform anterior process, triangular apically in lateral view; surstylus elliptical in lateral view |
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– | Arista pubescent (Fig. |
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7 | Mesonotum with 0 + 2 dorsocentral setae | 8 |
– | Mesonotum with 0 + 3 dorsocentral setae (exceptionally |
15 |
8 | Mesonotum with acrostichal setulae in six rows (exceptionally |
9 |
– | Mesonotum with acrostichal setulae in four rows | 13 |
9 | Acrostichal setulae with two pairs of strong setae in front of a pair of prescutellar seta |
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– | Acrostichal setulae with pair of strong setae in front of a pair of prescutellar setae | 10 |
10 | Arista plumose; only hind tarsomeres yellow |
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– | Arista pubescent; legs not as above | 11 |
11 | Surstylus long, extended part as long as height of epandrium, expanded apically in lateral view (Fig. |
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– | Surstylus short, extended part short than half height of epandrium, slender apically in lateral view | 12 |
12 | Syntergosternite 7+8 with one anterior median process in lateral view (Fig. |
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– | Syntergosternite 7+8 with one broad membranous process (Fig. |
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13 | Arista with microscopic setulae, shorter than 1/4 height of first flagellomere; male genitalia: surstylus narrow apically with falcate apical process in lateral view |
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– | Arista pubescent, with longest rays ca. 1/3 height of first flagellomere; male genitalia: surstylus wide apically with teeth or acute process in lateral view | 14 |
14 | Female sternite nine rectangular, ca. 3 × as wide as long, and sternite seven without triangular apical processes; male genitalia: surstylus with acute process projecting forwards in lateral view |
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– | Female sternite nine semicircular, sternite seven with pair of triangular apical processes; male genitalia: surstylus contorted claviform in lateral view and cone-shaped in posterior view, with two acute apical teeth |
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15 | Mesonotum with acrostichal setulae in two rows | 16 |
– | Mesonotum with acrostichal setulae in six rows | 17 |
16 | Anepisternum with bluish grey pruinescence; mid and hind tibiae yellow |
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– | Anepisternum with whitish grey pruinescence; mid and hind tibiae yellow except blackish apical 1/4 |
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17 | Mesonotum with brownish grey pruinescence, first postsutural dorsocentral setae weak, hairlike; male genitalia: surstylus fused with the epandrium, claviform with a triangular basal process, a projecting apical process, a small acute ventroapical process and a tiny incision in lateral view |
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– | Mesonotum with whitish grey pruinescence, first postsutural dorsocentral setae strong; male genitalia: surstylus articulated with epandrium, triangular or lobe-like with a single process | 18 |
18 | Face and parafacial with sparse whitish gray pruinescence; arista short plumose, longest rays slightly shorter than height of first flagellomere; mid femur with four anterior setae; male genitalia: epandrium with wide median incision on posterior margin in lateral view; surstylus short cone-shaped and originating from inner side of epandrium in lateral view and converging apically in posterior view; female unknown |
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– | Face with a yellow triangular median spot or only slightly yellow at middle of face, and parafacial with dense whitish gray pruinescence; arista with microscopic setulae, longest rays shorter than 1/3 height of first flagellomere; mid femur with three anterior setae; male genitalia: epandrium with small subapical incision in lateral view and acute apically in posterior view, surstylus contorted with apical setulae and originating from inner side of epandrium in lateral view; female sternite eight semicircular with pair of processes on anterior margin and a wide groove between processes |
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The specific epithet is named for holotype locality in Guizhou, Dashahe.
Arista blackish brown except paler basally. Mesonotum 1 + 3 dorsocentral setae, acrostichal setulae in four irregular rows, with two pairs of strong acrostichal setulae before prescutellar setae. Epandrium broad, ventral apical angle with setae. Surstylus separated from epandrium and extended from interior of epandrium, curved in lateral view, with connective sclerite under cercus in posterior view. Aedeagal apodeme long, nearly as long as aedeagus.
The new species is very similar to
China (Guizhou).
Latin,
Frons dark brown, anterior margin yellow, two black longitudinal stripes along fronto-orbital rows, extending from ocellar triangle to occiput. Arista blackish brown except paler basally, pubescent with longest setulae almost as long as 1/4 height of first flagellomere. Mesonotum with 0 + 2 dorsocentral setae, acrostichal setulae in six irregular rows. Surstylus separated from epandrium, distorted in lateral view and only one short curved claviform process, narrow apically and inwards curved in posterior view. Aedeagal apodeme long, nearly as long as aedeagus.
The new species is very similar to
China (Guizhou).
Latin,
Antenna brownish yellow, first flagellomere blackish brown on 2/3 apex of dorsal margin. Proboscis blackish brown, with pale yellow and black setulae. Mesonotum with two dorsocentral setae, acrostichal setulae in six irregular rows. Mid and hind tibia brownish yellow at base and tarsomeres dark yellow. Syntergosternite circular, broad dorsally and narrow ventrally, with one small anterior median process in lateral view, one membranous band under ventral bridge. Hypandrium median ventral side outward, V-shaped. Aedeagal apodeme long, slightly shorter than aedeagus.
The new species is very similar to
China (Yunnan).
The specific epithet is named for the holotype locality in Yunnan, Zhaotong.
Face and parafacial with dense grayish white pruinescence. Anterior fronto-orbital setae reclinate, shorter than posterior fronto-orbital setae. Mesonotum with 0+2 dorsocentral setae, acrostichal setulae in six irregular rows. Mid and hind tarsomeres dark yellow (sometimes fore tarsomeres also dark yellow). Wing slight yellow and hyaline, brown at base. Dorsal of epandrium narrower than dorsal of syntergosternite. Surstylus separated from epandrium, claviform, with single long seta in the middle, expanded apicially in lateral view and cone-shaped in posterior view. Postgonite hairy. Aedeagal apodeme tubular, as long as half-length of aedeagus.
The new species is very similar to
The new species is very similar to
China (Yunnan).
The key is modified from
modified from Shi and Yang 2014