Corresponding author: Ding Yang (
Academic editor: C. Borkent
Only seven species of the genus
Zhang B, Ren J, Li Y, Yang D (2020) New and little-known species of
The genus
The subgenus
So far, only the following seven species of the genus
So far, only the following seven species of the genus
The specimens were studied and illustrated with a ZEISS Stemi 2000-c stereomicroscope. Details of coloration were checked in specimens immersed in 75% ethyl alcohol (C2H5OH). Genitalic preparations of males were made by macerating the apical portion of the abdomen in cold 10% NaOH for 12–15 hours. After examination, it was transferred to fresh glycerine (C3H8O3) and stored in a microvial pinned below the specimen. The specimens studied, which were collected in Laos during June 2017, are deposited in the
Some type and non-type material used in this paper were borrowed from the
The morphological terminology mainly follows
1 | Body velvet black or velvet orange (Figs |
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– | Body brownish yellow or brown (Figs |
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2 | Thorax velvet black ( |
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– | Thorax velvet orange (Figs |
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3 | Clasper of gonostylus with two short acute blackened spines ( |
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– | Clasper of gonostylus complex, beak with a lump near tip, back of beak bearing a sickle-shaped prominence directed anteriorly; dististyle outer surface with an acute blackened spine curved up (Fig. |
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4 | Clasper of gonostylus broad and short, beak slender ( |
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– | Clasper of gonostylus slender, beak short ( |
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5 | Body large, greater than 20 mm; eighth sternite extended backward, posterior margin with deep median notch, two small triangular processes present at bottom of notch (Fig. |
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– | Body not exceeding 20 mm; eighth sternite not as above |
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6 | Posterior margin of cell cua1 as wide as base (Figs |
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– | Posterior margin of cell cua1 narrower than base (Fig. |
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7 | Rs a little longer than R2+3, relatively straight (Fig. |
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– | Rs very short, equaling R2, which is distinct and oblique ( |
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Hypopygium is blackish with black setae. Posterior margin of ninth tergite has a low U-shaped notch. Posterior margin of eighth sternite has a pair of digitiform appendages. Clasper of gonostylus is complex, broad, and short; beak slender.
Male (
Head (Figs
Thorax (Figs
Abdomen (Fig.
Hypopygium (Figs
Female (
Female resembles male in head and thorax, except abdomen plump. Eighth tergite and eighth sternite black throughout with black setae. Ninth tergite, ninth sternite, and tenth tergite dark black, with black setae (Fig.
Ovipositor (Figs
3 males 1 female (
China (Guizhou, Yunnan), Laos (Houaphanh).
Hypopygium covered with dense long setae. Clasper of the gonostylus slender; beak short with a long tuber which directed upward; dorsal margin bearing a sickle-shaped prominence which directed anteriorly (
Recorded from Nepal and Upper Burma (
India (Assam), Laos, Myanmar, Nepal.
Abdomen yellowish-white pruinose has blackish brown stripes, last two segments dark brown. Male ninth tergite has a slender hammer-shaped projection at middle. Clasper of the gonostylus with two short acute blackened spines (
Paratype, male, Laos: Nam Mat, 15 April 1918 (
China (Yunnan), Laos.
Abdomen is reddish brown with thick brownish yellow setae. Eighth sternite extended backward, posterior margin has a deep median notch. Posterior margin of ninth tergite extended with sclerotized depression. Lobe of gonostylus is fleshy with a V-shaped depression at posterior margin. Clasper of gonostylus is duck-shaped.
Male (
Head (Figs
Thorax (Figs
Abdomen (Fig.
Hypopygium (Figs
Semen pump (Figs
Female (
Female resembles male in head and thorax, except abdomen plump (Fig.
Ovipositor reddish brown (Figs
Laos (Champasak).
The species is named after Champasak Province, where the type locality is located.
This new species is somewhat similar to
Thoracic prescutum has three brown stripes; wing is brownish yellow and with brown pterostigma; Rs is a little longer than R2+3 and relatively straight; petiole of cell m1 is short. Abdomen is brownish yellow; notch of ninth sternite has a depressed semicircular lobe; lobe of gonostylus slender and gradually narrowed to obtuse tip, before the apex slightly narrower; clasper of gonostylus is a shallow, beak blackened, surface with abundant minute setae, lower lobe well developed (
Holotype, male, India: Kumaon, Muktesar, Khasi Hills, Cherrapunji, 1 April 1922 (
China (Xizang), India (Assam, Uttarakhand, Uttar Pradesh, W Bengal), “Indochina” (= ?Laos), Tajikistan.
Rs is relatively long and cell m1 is petiolate. Tip of eighth sternite has long dense thick setae. Ninth tergite has a U-shaped depression. Lobe of gonostylus is subtriangular and posterior margin has a shallow V-shaped incision with a black sclerotized protuberance. Clasper of gonostylus is small and beak-like.
Male (
Head (Figs
Thorax (Figs
Abdomen (Fig.
Hypopygium (Figs
Semen pump (Figs
3 males (
China (Chongqing), India (Assam), Laos (Tong La, Houaphanh), Nepal, Thailand (Chiang Mai).
We are grateful to Dr Liang Wang (Beijing) for collecting the specimens from Laos. We are also particularly grateful to Dr Zacariah Billingham, Dr Matthew J. Petersen, and Dr Chris Borkent for their valuable suggestions on this paper. The research was funded by the National Natural Science Foundation of China (no. 31501880) and the Ministry of Science and Technology of the People’s Republic of China (2005DKA21402).