Corresponding author: Boutaïna Belqat (
Academic editor: V. Blagoderov
A total of 54 species of
Banamar O, Chandler PJ, Driauach O, Belqat B (2020) New faunistic records of the family Mycetophilidae (Insecta, Diptera) from Morocco. ZooKeys 934: 93–110.
The family
The fungus gnat fauna of Morocco is practically unknown. The first information on Moroccan
Five species, without given localities, were recorded by
Three other species were reported by
Recent records were added by
This paper is the first contribution relating specifically to the
A total of 724 specimens of
Most of the material was collected by O. Banamar, O. Driauach, and B. Belqat. Additional material was provided by Dr M. Ebejer that he had collected in Morocco and some Moroccan
A list of sampling sites, with coordinates and altitudes, is given in Table
A first checklist of Moroccan
Map showing all collecting localities for
Sampling sites (in alphabetical order) harbouring the species collected in Morocco, in the present study, with localities, geographical coordinates and altitudes.
Province | Station | Locality | Elevation (m) | Geographical coordinates |
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Tétouan | Marabout El Khaloua | Dar Khennouss | 788 |
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Tanger | Grotte d’Hercule | Tanger | 12 |
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Chefchaouen | Aïn El Ma Bared | Bouzthate | 1267 |
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Aïn El Malâab | Parc National Talassemtane | 1278 |
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Aïn Ras El Ma | Majjou | 856 |
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Aïn Sidi Brahim Ben Arrif | Larache | 897 |
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Aïn Takhninjoute | Bab Rouida | 1512 |
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Aïn Tiouila | Parc National Talassemtane | 1502 |
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Bab El Karne | Tamakoute | 1248 |
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Cascade Chrafate | Chrafate | 820 |
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Daya Afersiw | Mezine | 303 |
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Daya Amsemlil | Jbel Bouhachem | 1059 |
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Daya avant Taïda | Taïda | 425 |
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Daya Fifi | Fifi | 1202 |
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Daya Jbel Zemzem | Jbel Zemzem | 216 |
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Daya Mtahen | Jbel Bouhachem | 966 |
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Daya Tazia | Route Moulay Abdessalam | 721 |
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Douar Abou Boubnar (Marabout Sidi Gile) | Parc National Talassemtane | 1247 |
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Douar Kitane | Kitane | 52 |
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Douar Tizga | Amsa | 516 |
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Forêt-Adrou | Parc National Bouhachem | 580 |
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Forêt-Aïn Boughaba | Jbel Bou Bessoui | 1526 |
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Forêt-Jbel Lekraa | Parc National Talassemtane | 1541 |
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Forêt-Taghzoute | Parc National Bouhachem | 556 |
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Maison forestière | Parc National Talassemtane | 1674 |
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Oued Aârate | Dardara | 269 |
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Oued Amsemlil | Jbel Bouhachem | 1059 |
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Oued à 15 Km de Fifi | Bouzthate | 1256 |
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Oued Kelâa | Akchour | 303 |
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Oued Majjou | Majjou Village | 799 |
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Oued Majjou (Hafa Meqlouba) | Chefchaouen | 825 |
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Oued Sidi Yahya Aârab | Sidi Yahya Aârab | 62 |
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Oued Tisgris | Hmmadesh | 490 |
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Oued Tkarâa | Jbel Bouhachem | 959 |
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Route Ksar El Kébir-Chefchaouen | Tattofte | 133 |
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Berkane | Grotte des Pigeons | Beni Snassen | 676 |
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Grotte du Chameau | Beni Snassen | 427 |
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Oued Tafoughalt | Tafoughalt | 751 |
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Meknès | Aïn Walili | Sidi Moulay Idriss Zerhoun | 490 |
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Ifrane | Forêt-3.5 km S. Azrou | Azrou | 1450 |
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Cited from North Africa (Tunisia) by
Rif: Daya Tazia, 1♂ 12/V/2015.
Rif: Oued Aârate, 1♂, 26/III/2014.
Cited from North Africa by
Rif: Forêt-Jbel Lekraa, 1♂, 12/VI/2013, coll. Ebejer; Oued Tisgris, 1♂, 25/III/2014; Oued Aârate, 1♂, 26/III/2014; Daya Tazia, 1♂, 25/IV/2014; Oued Majjou (Majjou Village), 2♂♂, 10/V/2014; Douar Tizga, 1♂, 25/VI/2014; Maison forestière, 2♂♂, 5♀♀, 07/VI-17/VI/2014, Malaise trap; Daya Jbel Zemzem, 1♂, 2/II/2015; Aïn Takhninjoute, 1♂, 21/IV/2015; Forêt-Adrou, 1♂, 25/IV/2015, coll. Ebejer; Daya Amsemlil, 2♂♂, 23/IV/2016.
Rif: Daya Fifi, 1♂, 23/XI/2012; Oued Aârate, 1♂, 26/III/2014, 2♂♂, 07/V/2015; Daya Tazia, 1♂, 25/IV/2014; Oued Majjou (Majjou Village), 3♂♂, 7♀♀, 10/V/2014; Aïn El Malâab, 2♂♂, 2♀♀, 17/V/2014; Aïn El Malâab, 12♂♂, 17/V/2014; Maison forestière, 1♂7♀♀, 18/V/2014, 6♂♂, 17/VI/2014; Douar Tizga, 1♂, 25/VI/2014; Aïn Takhninjoute, 1♂, 21/IV/2015; Daya Amsemlil 1♂, 2♀♀, 23/IV/2016; Daya Mtahen, 1♂, 16/V/2016; Daya avant Taïda, 1♂, 20/IV/2018.
Recorded from North Africa (Tunisia) (
Rif: Chefchaouen, VII/1975, 1♂, coll. J. Beaucournu (
Recorded from North Africa (Tunisia) (
Rif: Daya Afersiw, 1♂, 11/VI/2013; Forêt-Jbel Lekraa, 1♂, 12/VI/2013, coll. Ebejer.
Recorded from Morocco (
Rif: Douar Abou Boubnar, 1♂, 18/V/2014; Maison forestière, 6♂♂, 1♀, 17/VI/2014, Forêt-Adrou, 1♂, 25/IV/2015, coll. Ebejer; Oued Sidi Yahya Aârab, 1♂, 1♀, 25/IV/2015, coll. Ebejer. Beni Snassen: Oued Tafoughalt, 1♂, 25/XII/2014.
Rif: Oued Majjou (Majjou Village), 1♂, 09/IV/2013.
Rif: Forêt-Aïn Boughaba, 1♂, 24/V/2013.
Rif: Maison forestière, 1♂, 4/VII/2013, 12♂♂, 17/VI/2014, 1♂, 17/III/2015; Oued Tkarâa, 1♂, 1♀, 16/V/2016; Oued Sidi Yahya Aârab, 1♂, 25/IV/2015.
Rif: Oued Kelâa, 1♀, 13/II/2013; Oued Aârate, 1♀, 26/III/2014. Beni Snassen: Grotte des pigeons, 1♂, 25/XII/2014; Oued Tafoughalt, 1♂, 25/XII/2014.
Rif: Bab El Karne, 1♂, 25/XII/2016.
Cited from Morocco: High Atlas, Toubkal (
Rif: 20 km west of Targuist, Ketama, 23/IV/1966, 6♂♂, 3♀♀, coll. A.M. Hutson (
Cited from North Africa (Tunisia) (
Rif: Oued Kelâa, 2♂♂, 13/II/2013; Forêt-Jbel Lekraa, 1♂, 12/VI/2013, coll. Ebejer; Douar Kitane, 1♂, 01/I/2015; Daya Amsemlil, 1♂, 23/IV/2016.
Cited from North Africa (Algeria) (
Rif: Cascade Chrafate, 1♂, 1♀, 28/IV/2015.
Rif: Source Aheramen, 1♂, 10/V/2014.
Cited from North Africa (Algeria) (
Rif: Maison forestière, 1♀, 17/VI/2014; Daya Amsemlil, 1♂, 28/II/2015; Aïn Tiouila, 1♂, 02/V/2015; Daya Fifi, 1♂, 25/XII/2015.
Cited from Morocco (
Beni Snassen: Grotte des pigeons, 1♂, 25/XI/2014.
Cited from Morocco (
Rif: Chefchaouen, VII/1975, 1♀, coll. J. Beaucournu,
Cited from North Africa (Algeria) (
Rif: Maison forestière, 1♀, 18/V/2014; Daya Amsemlil, 1♂, 28/II/2015; Cascade Chrafate, 1♂, 1♀, 28/IV/2015; Daya Amsemlil, 1♀, 23/IV/2016. Grotte-Aïn El-Aouda, 1♂, 1♀, 2/V/1914,
Rif: Chefchaouen, VII/1975, 13♂♂, 6♀♀, coll. J. Beaucournu,
Recorded from Morocco (
Rif: Forêt-Aïn Boughaba, 2♂♂, 24/V/2013; Forêt-Jbel Lekraa, 2♂♂, 1♀, 12/VI/2013, coll. Ebejer; Douar Kitane, 1♂, 13/III/2014, Malaise trap; Aïn El Malâab, 11♂♂, 17-V-2014; Aïn Takhninjoute, 1♂, 17/V/2014; Oued à 15 km de Fifi, 2♂♂, 06/V/2014; Aïn El Ma Bared, 3♀♀, 06/V/2014; Maison forestière, 2♀♀, 18/V/2014; Oued Majjou (Hafa Meqlouba), 1♂, 10/V/2014; Oued Majjou (Majjou Village), 1♂, 10/V/2014; Aïn Takhninjoute, 1♂, 17/V/2014; Maison forestière, 2♀♀, 17/VI/2014; Grotte d’Hercule, 3♂♂, 2♀♀, 29/III/2015; Aïn Takhninjoute, 1♂, 21/IV/2015, 1♂, 24/VI/2015; Oued Aârate, 1♂, 07/V/2015; Bab El Karne, 1♂, 25/XII/2015; Daya Amsemlil, 1♀, 1♂, 23/IV/2016; Daya Mtahen, 1♂, 16/V/2016; Daya avant Taïda, 1♀, 20/IV/2018; Forêt-Taghzoute, 1♂1♀, 25/IV/2015, coll. Ebejer.
Forêt-Taghzoute, 1♂, 25/IV/2015; Route Ksar El Kébir-Chefchaouen, 2♂♂, 5/VI/2013, coll. Ebejer.
Rif: Daya Fifi, 2♂♂, 26/III/2014; Daya Tazia, 1♀, 25/IV/2014; Oued Tkarâa, 1♀, 16/V/2016; Forêt-Jbel Lekraa, 1♀, 12/VI/2013, coll. Ebejer; Forêt-Taghzoute, 1♀, 25/IV/2015, coll. Ebejer. Beni Snassen: Grotte d’Hercule, 1♀, 29/III/2015.
Rif: Forêt-Jbel Lekraa, 1♂, 12/VI/2013, coll. Ebejer; Oued Amsemlil, 1♂, 28/II/2015, Daya Amsemlil, 1♂, 25/XII/2015; Daya Amsemlil, 2♂♂, 23/IV/2016; Daya avant Taïda, 1♂, 20/IV/2018.
Rif: Chefchaouen, VII/1975, 4♂♂, coll. J. Beaucournu,
Cited from North Africa (
Rif: Forêt-Aïn Boughaba, 1♂, 1♀, 24/V/2013; Aïn El Malâab, 1♂, 14/VII/2013; Maison forestière, 2♂♂, 17/VI/2014; Bab El Karne, 1♂, 25/XII/2015; Daya Amsemlil, 3♂♂, 1♀, 23/IV/2016; Douar Kitane, 5♂♂, 1♀, 29/XII/2016.
Cited from Morocco (
Rif: Forêt-Jbel Lekraa, 1♀, 12/VI/2013, coll. Ebejer; Oued Majjou (Majjou Village), 1♂, 10/V/2014.
Cited from Morocco (
Rif: Forêt-Jbel Lekraa, 1♂, 12/VI/2013, coll. Ebejer.
Rif: Forêt-Jbel Lekraa, 1♂, 12/VI/2013, coll. Ebejer; Forêt-Aïn Boughaba, 1♂, 24/V/2013; Oued à 15 km de Fifi, 1♂, 06/V/2014; Aïn El Malâab, 1♂, 17/VI/2014.
Cited from North Africa (Tunisia) (
Rif: Forêt-Taghzoute, 1♂, 25/IV/2015, coll. Ebejer; Oued Aârate, 1♂, 26/III/2014.
Rif: Oued Kelâa, 1♂, 13/II/2013.
Rif: Forêt-Jbel Lekraa, 2♂♂, 12/VI/2013, coll. Ebejer; Maison forestière, 1♂, 1♀, 18/V/2014, Daya Amsemlil, 7♂♂, 25/IV/2016.
Recorded from Morocco (
Rif: Daya Tazia, 1♂, 25/IV/2014; Aïn El Ma Bared, 1♀, 06/V/2014; Aïn El Malâab, 1♀, 17/V/2014; Aïn Takhninjoute, 1♂, 17/V/2014, 1♀, 21/IV/2015; Maison forestière, 1♂, 17/VI/2014; Grotte d’Hercule, 1♂, 29/III/2015; Daya Amsemlil, 4♂♂, 3♀♀, 23/IV/2016.
Rif: Daya Amsemlil, 1♂, 28/II/2015.
Rif: Daya Afersiw, 1♂, 11/VI/2013; Oued Aârate, 1♂, 26/III/2014.
Recorded from North Africa (Tunisia and Algeria) (
Rif: Oued Kelâa, 1♂, 1♀, 13/II/2013; Oued Aârate, 1♂, 3♀♀, 26/III/2014, 1♂, 07/V/2015; Oued Majjou (Majjou Village), 1♂, 10/V/2014; Aïn Takhninjoute, 2♂♂, 17/V/2014; Grotte d’Hercule, 1♂, 29/III/2015.
Rif: Forêt-Jbel Lekraa, 1♂, 12/VI/2013, coll. Ebejer; Maison forestière, 1♂, 18/V/2014; Daya Amsemlil, 2♂♂, 28/II/2015; Aïn Takhninjoute, 6♂♂, 5♀♀, 21/IV/2015; Daya Amsemlil, 3♂♂, 1♀, 23/IV/2016.
Rif: Forêt-Jbel Lekraa, 1♂, 12/VI/2013, coll. Ebejer; Oued Aârate, 1♂, 26/III/2014; Maison forestière, 1♂, 1♀, 18/V/2014; Daya Amsemlil, 3♂♂, 3♀♀, 28/II/2015, 1♂, 1♀, 23/IV/2016; Cascade Chrafate, 1♂, 28/IV/2015; Bab El Karne, 14♂♂, 1♀, 25/XII/2015.
Rif: Maison forestière, 1♂, 17/VI/2014; Aïn El Malâab, 1♂, 17/V/2014.
Rif: Oued à 15 km de Fifi, 2♂♂, 06/V/2014; Oued Aârate, 1♂, 07/V/2015.
Rif: Aïn Takhninjoute, 1♂, 21/IV/2015; Bab El Karne, 3♂♂, 1♀, 25/XII/2015; Daya Amsemlil, 1♂, 23/IV/2016; Oued Tkarâa, 1♂, 16/V/2016.
Beni Snassen: Grotte du chameau, 1♂, 24/XI/2015.
Recorded from North Africa (Algeria) (
Rif: Forêt-Jbel Lekraa, 1♀, 12/VI/2013, coll. Ebejer; Bab El Karne, 1♂, 25/XII/2015.
Beni Snassen: Grotte du Chameau, 1♂, 24/XI/2015.
Rif: Maison forestière, 1♂, 17/VI/2014.
Rif: Aïn Ras El Ma, 1♂, 1♀, 3/V/2013; Oued Aârate, 1♂, 26/III/2014.
Rif: Forêt-Jbel Lekraa, 1♂, 12/VI/2013, coll. Ebejer; Maison forestière, 1♂, 07/VI-17/X/2014, Malaise trap; Maison forestière, 1♂, 17/VI/2014; Aïn Takhninjoute, 1♂, 21/IV/2015; Daya Amsemlil, 1♀, 23/IV/2016.
Cited from North Africa (Tunisia) (
Rif: Oued Majjou (Majjou Village), 1♂, 09/IV/2013; Oued Sidi Yahya Aârab, 1♀, 25/IV/2015. Middle Atlas: Aïn Walili, 1♂, 18/II/2016.
Cited from North Africa (Algeria) (
Rif: Aïn Ras El Ma, 1♀, 27/III/2013.
Cited from Morocco (
Rif: Daya Tazia, 1♂, 25/IV/2014; Maison forestière, 1♂, 07/VI-17/X/2014, Malaise trap; Aïn El Ma Bared, 1♂, 25/XII/2015. Middle Atlas: Forêt-3.5 km S. Azrou, 1♀, 8/V/2012, coll. Ebejer.
Cited from North Africa (Tunisia) (
Rif: Maison forestière, 10♂♂, 1♀, 07/VI-17/VI/2014, 1♂, 17/XI/2015, Malaise trap.
Rif: Oued Aârate, 1♂, 26/III/2014; Maison forestière, 11♂♂, 07/VI-17/VI/2014, 9♂♂, 17/XI/2015, Malaise trap.
Rif: Daya Fifi, 3♂♂, 23/XI/2012; Oued Aârate, 1♂, 26/III/2014; Daya Fifi, 1♂, 26/III/2014; Aïn Sidi Brahim Ben Arrif, 1♂, 25/IV/2014; Daya Amsemlil, 2♂♂, 28/II/2015; Grotte d’Hercule, 1♂, 29/III/2015, Cascade Chrafate, 2♂♂, 28/IV/2015; Oued Sidi Yahya Aârab, 1♂, 25/IV/2015; Aïn El Ma Bared, 2♂♂, 25/XII/2015; Bab El Karne, 2♀♀, 25/XII/2015. Middle Atlas: Forêt-3.5 km S. Azrou, 2♂♂, 8/V/2012, coll. Ebejer.
Rif: Daya Fifi, 1♀, 23/XI/2012; Oued Kelâa, 1♂, 13/II/2013; Daya Fifi, 1♂, 26/III/2014; Oued Amsemlil, 2♂♂, 2♀♀, 28/II/2015, 1♂, 31/I/2017; Aïn Takhninjoute, 1♂, 21/IV/2015; Cascade Chrafate, 2♂♂, 28/IV/2015; Bab El Karne, 1♀, 25/XII/2015; Daya Amsemlil, 1♀, 23/IV/2016; Daya avant Taïda, 2♂♂, 20/IV/2018.
Rif: Aïn Sidi Brahim Ben Arrif, 1♂, 25/IV/2014; Daya Tazia, 1♂, 25/IV/2014; Forêt-Adrou, 1♀, 25/IV/2015, coll. Ebejer; Daya Amsemlil, 1♂, 23/IV/2016; Daya avant Taïda, 1♂, 20/IV/2018.
Rif: Daya Amsemlil, 1♂, 23/IV/2016.
Rif: Forêt-Aïn Boughaba, 1♂, 24/V/2013. Beni Snassen: Oued Tafoughalt, 1♂, 25/XII/2014.
Rif: Aïn El Malâab, 1♂, 17/V/2014; Maison forestière, 40♂♂, 2♀♀, 07/VI-17/VI/2014, Malaise trap.
Rif: Oued Amsemlil, 6♂♂, 1♀, 28/II/2015; Aïn Sidi Brahim Ben Arrif, 2♂♂, 25/IV/2014.
Rif: Daya Fifi, 2♂♂, 26/III/2014.
Cited from North Africa (Algeria) (
Rif: Oued Majjou (Majjou Village), 1♂, 09/IV/2013; Maison forestière, 46♂♂, 1♀, 17/VI/2014.
Cited from Morocco: Rabat, Oued y Kern, V/1973, coll. H. Choumara,
Rif: Maison forestière, 1♂, 4/VII/2013, 4♂♂, 2♀♀, 07/VI-17/X/2014, Malaise trap, 1♂, 1♀, 17/VI/2014; Daya Jbel Zemzem, 1♂, 02/III/2015; Oued Sidi Yahya Aârab, 1♂, 25/IV/2015; Bab El Karne, 1♂, 25/XII/2015; Marabout El Khaloua, 1♂, 03/VI/2018.
The majority of the species recorded here are widespread in the Mediterranean region and also more widely in Europe and the Palaearctic Region. In addition to these, 10 species considered to be new to science were also found, and these will be treated elsewhere. These new species belong to the genera
Knowledge of the North African
The
Moroccan species whose distribution is entirely or principally Mediterranean are the following, with the distribution outside Morocco stated in each case:
We are grateful to Dr Martin Ebejer who kindly provided material that he collected in Morocco and for permitting us to publish his new records. We also thank the authorities of