Corresponding author: Remko Leijs (
Academic editor: T. Dörfel
The species in the subgenus
Leijs R, Dorey J, Hogendoorn K (2020) The genus Amegilla (Hymenoptera, Apidae, Anthophorini) in Australia: a revision of the subgenus Asaropoda. ZooKeys 908: 45–122.
The Australian members of the genus
The taxonomy of the genus
The first
Within the subgenus
Here we present a revision of the species in the subgenera
This study is based on examination of 1466 museum specimens, including most type specimens. The following acronyms are used in the database of the examined specimens (available as supplementary information associated with this paper):
Additional fresh specimens were collected throughout Australia by the authors and other collectors acknowledged below. Field collected specimens were killed and preserved in absolute ethanol to allow DNA extraction at a later stage. Ethanol preserved specimens, as well as extracted DNA, are kept in the Australian Biological Tissue Collection (
Genitalia were dissected from male museum specimens, after relaxing in a humid container with added chromocresol as fungicide for up to three days. Genitalia were treated with 10% cold sodium hydroxide for 24 hours, glacial acetic acid for approximately one hour and stored in glycerol to facilitate the study of morphology. For species descriptions the terminology used by
A Leica stereomicroscope with auto-montage imaging stacking software was used to obtain high-resolution diagnostic images of all species. A Canon EOS 5DSR camera attached to a Nikon Eclipse 50i compound microscope and Zerene Stacker were used to image male genitalia and metasomal sterna seven and eight.
The following abbreviations are used in the identification key and species descriptions:
DNA extraction, PCR amplification and sequencing were performed as described in
Some of the specimens treated here were also submitted to BOLD (Barcode of Life Database) for DNA barcoding using the cytochrome c oxidase subunit 1 gene. Specimen details, including DNA sequence, collection dates and locality information can be accessed in BOLD under the project Australian Bee Survey, e.g.,
Phylogenetic analyses of aligned sequence data were carried out using PAUP* version 4.0b8 (
MRBAYES allowed the application of relaxed molecular clock methods in order to obtain estimates of node divergence times. Because fossils are unavailable for
Morphological examination of type specimens in Australia and abroad and specimens from all major Australian museum collections, as well as newly collected material during Bush Blitz surveys and other collecting trips by the authors, revealed 21 species of which ten were recognised as new, while 15 names needed to be synonymised. Some of the male types of species described by
The main diagnostic characters for
DNA data were generated for 12 species, including six of the new species. These species were analysed together with data from other Australian
Bayesian molecular phylogenetic consensus tree of CO1 sequences. Node support values (posterior probabilities) are shown near the nodes. The terminals are labelled with species names (new species in red) and RB-numbers, which refer to Table S1.
The good match between the molecular and morphological data types allows the allocation of species for which no DNA data are as yet available into species groups. The differences in morphology between these groups, which are discussed below, are more pronounced than those between
The
The
The
The
The three remaining species in the phylogenetic tree may contain the following lineages:
The
The
The
It is difficult to place
Considering the good match between phylogenetic species groups and their morphology, which mainly are male sexual characters, it is interesting to note that there are large dissimilarities in general appearance, mostly in pubescence colouration, between closely related species and this is especially the case when closely related species have overlapping or nearly overlapping distributions. Compare for example
General distribution patterns of individual species of
Often, closely related species do not overlap in distribution. For example, there are three cases of sister species pairs that are distributed east and west of the Gulf of Carpentaria in Northern Australia:
While for several species an accurate understanding of the phenology is hampered by small sample sizes, some general patterns can be deduced. As in other
From the following
It is remarkable that for
The diagnosis for this subgenus as given by
In the following text, the main subgeneric diagnostic characters are in bold:
We now recognise 21 species in
1 | Forewing: hairs absent in 3rd submarginal cell and 2nd medial cell, usually also in 2nd submarginal cell and 1st medial cell; S4 in males with a rounded, triangular or elongated brush of dense usually forward-directed bristles, S6 of females medially with raised area which varies among species from inconspicuously roughened, broadly parabolic with well-defined rim and slender spine, to almost a pygidium like structure |
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– | Forewing: hairs present in in 3rd submarginal cell, 2nd medial cell and most other cells; S4 in males and S6 in females not modified as described above |
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2 | Female: Integument of paraclypeal areas black; fore and mid femora and tibiae with iridescent blue-green hairs, male: S6 gently convex; apex S7 broadly triangular |
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– | Female: Integument of paraclypeal areas partly yellow, white or ivory; hair on fore and mid legs never iridescent, male: S6 with broad depressions either side of midline; apex S7 ovate |
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1 | Female: Body size large (> 19 mm), pubescence on head and mesosoma white, metasomal integument predominantly dark red (Fig. |
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– | Not with above combination of characters (20 spp.) |
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2 | Metasomal pubescence predominantly black, some species with light coloured posterior bands of white pubescence on terminal terga (5 spp.) |
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– | Metasomal pubescence predominantly orange, brown or grey, with or without an anterior band of black pubescence on T2 (15 spp.) |
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3 | Light coloured posterior bands present on all terga (Figs |
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– | Light coloured posterior bands lacking, or only on last 1-3 terminal terga (Figs |
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4 | Female: T1 with thin, faint white posterior band, T2-4 with well-developed white posterior hair bands (Fig. |
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– | All terga with well-developed posterior hair bands. Hair bands on T1-2 orange (2 spp.) |
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5 | Female: clypeus ivory with two large black marks, which are sometimes combined at the top, scape black. Male: S4 with two adjacent small patches of downward-directed bristles on apicomedial margin and two large comma-shaped patches of bristles on S5 (Fig. |
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– | Female: clypeus with two small brown marks, scape with pale yellow mark (Fig. |
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6 | Metasomal pubescence entirely black (Fig. |
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– | T4-5 (female), T5-6 (male) with predominantly white hairs (Fig. |
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7 | Male |
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– | Female |
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8 | Both S3 and S4 medially with round patch of bristles, the patch on S3 smaller than on S4 (Figs |
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– | Only S4 with median patch of bristles of varying size and shape (Figs |
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9 | S2-4 without apicomedial emarginations; S5 apicomedial area deeply emarginated, emargination deeper than wide and almost parallel to body axis (Figs |
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– | S2-4 with distinct apicomedial emarginations (Fig. |
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10 | Apicomedial emargination on S5 parallel-sided, slightly deeper than wide (Fig. |
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– | Apicomedial emargination on S5 more than twice as deep as wide (Fig. |
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11 | S4 shape of patch of bristles round (Figs |
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– | S4 shape of patch of bristles triangular or bell-shaped (Figs |
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– | S4 shape of patch of bristles elongated (Figs |
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12 | S4 patch of bristles relatively large, circa 1/7 of width of sternum (Figs |
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– | S4 patch of bristles small, less than 1/10 of width of sternum (Fig. |
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13 | Face marks pale yellow, scape brown (Fig. |
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– | Face marks ivory, underside of scape with ivory mark (Fig. |
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14 | S4 patch of bristles small, circa 1/5 of width of sternum (Fig. |
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– | S4 patch of bristles larger, S5 apicomedial emargination shallow (Figs |
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15 | S4 patch of bristles large, circa 1/3 of width of sternum, bristles brown (Fig. |
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– | S4 patch of bristles less than 1/3 of width of sternum, bristles black (2 spp.) |
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16 | S4 patch of bristles triangular (Fig. |
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– | S4 patch of bristles bell-shaped (Fig. |
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17 | S4 patch of black bristles broad and circa 1/2-2/3 of sternum width (Fig. |
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– | S4 patch of pale bristles narrow, circa 1/3 of width of sternum, S6 apicomedial area without conspicuous fimbria of black and orange-fringed hairs, T2 anteriorly with area of black hairs much narrower than marginal zone, or without black hairs (4 spp.) |
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18 | Face marks ivory/white, elongated patch of bristles on S4 black (Fig. |
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– | Face marks pale to bright yellow, patch of bristles on S4 dark or light brown (3 spp.) |
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19 | T2 anteriorly without dark hairs, S5 apicomedial emargination circa 6 × as wide as deep, (Fig. |
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– | T2 anteriorly with entire narrow band of dark hairs, S5 apicomedial emargination less than 4 × as wide as deep |
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20 | S6 marginal zone smooth without punctures or hairs, some short hairs medially (Fig. |
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– | S6 marginal area, smooth with open punctures and hairs, medially with patch of stiff dark hairs (Fig. |
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21 | Hind leg: tibial scopa on outer surface with white or grey-white hairs (5 spp.) |
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– | Hind leg: tibial scopa on outer surface with pale yellow, ochre or orange hairs (8 spp.) |
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22 | Paraclypeal area with pale yellow mark (3 spp.) |
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– | Paraclypeal area black (3 spp.) |
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23 | Paraclypeal marks small (Fig. |
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– | Paraclypeal marks large (Figs |
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24 | Scutum and T1 pubescence light brown, T2 and T3 anteriorly with black hairs |
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– | Scutum and T1 pubescence appears grey, hairs light ochre intermixed with longer black hairs, only T2 with black hairs anteriorly |
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25 | Supraclypeal area black, clypeus with yellow inverted T shaped mark (Fig. |
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– | Supraclypeal area with yellow or pale yellow mark, clypeus with two vertical brown marks (Fig. |
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26 | Paraclypeal area with light coloured mark (3 spp.) |
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– | Paraclypeal area without light coloured mark (6 spp.) |
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27 | T2 anteriorly with narrow band of black hairs across whole width, sterna with dark hairs |
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– | Dark hairs on T2 restricted to antero-lateral corners or lacking, sterna on disk with orange hairs (2 spp.) |
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28 | Integument of T3 and following segments anteriorly dark brown, clypeus with two well defined vertical dark orange marks (Fig. |
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– | Integument of T3 and following segments anteriorly orange, marks on clypeus undefined and small (Fig. |
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29 | T2 anteriorly with black hairs (2 spp.) |
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– | Pubescence on T2 uniformly coloured (4 spp.) |
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30 | Scape orange, T2 anterior with wide area of black hairs, clypeus marks undefined (Fig. |
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– | Scape black, apically brown, T2 anteriorly with narrow area of erect black hairs, clypeus with brown horseshoe-like mark (Fig. |
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31 | Clypeus colouration yellow, orange pubescence on terga with iridescent shine, S6 projection broadly parabolic, well defined (Figs |
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– | Clypeus colouration pale yellow or ivory, pubescence on terga without iridescent shine, S6 projection rounded, not well defined (Figs |
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32 | Clypeus marked with inverted yellow T (Fig. |
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– | Clypeus yellow, with two faint brown patches (Fig. |
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33 | Clypeus with pale yellow inverted T-shape, supraclypeal area black (Fig. |
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– | Clypeus colouration ivory, supraclypeal area black with narrow ivory line at base (Fig. |
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(22 males, 5 females).
Holotype of
Holotype of
Holotype of
Holotype of
The emargination and hair patches of the sterna of the males of
Male with triangular patch of black bristles on S4 and shallow emarginated apicomedial area of S5. Female with narrow yellow paraclypeal mark along clypeus, supraclypeal mark present, tibial scopa orange, T2 with black hairs anteriorly, process on S6 irregularly striate, not well defined posteriorly (Fig.
Male (AM K470111): Body length 14 mm, forewing length 9.7 mm, head width 5.0 mm.
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No. of records: | 4 | 12 | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 1 | 0 |
Figure
(7 males, 2 females).
Holotype, male, Useless loop road, 8-9 km from Derham Road, WA (
Allotype, female, WAM 21904, same locality data as holotype;
Paratype, male, WAM 21906, same locality data as holotype.
Face marks of both sexes ivory. Male S4 with rectangular shaped patch of black bristles, and broadly emarginated posterior rim of S5. Female paraclypeal and supraclypeal marks absent, tibial scopa outer surface orange, T2 with a few black hairs in anterolateral corners, process on S6 striate, little defined posteriorly (Fig.
Male holotype (WAM21905): Body length 14 mm, forewing length 9.2 mm, head width 5.0 mm.
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No. of records: | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 9 | 0 |
Figure
The specific epithet refers to the white face marks of this species.
17 males, 12 females.
Holotype of
Male with almost triangular patch of light brown stiff bristles on S4 and emarginate apicomedial area of S5 surrounded by dense branched hairs, lateral corners of S5 protruding. Female paraclypeal and supraclypeal marks present, tibial scopa ochre, T2 with a few black hairs anteriorly, process on S6 large narrow, posteriorly with angular smooth ridge Fig.
Male (SAMA 32-033598, RL0456) Body length 14 mm, forewing length 10.1 mm, head width 4.7 mm.
redescription (VM 35933): Body length 14.5 mm, forewing length 10.6 mm, head width 5.2 mm.
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No. of records: | 1 | 0 | 1 | 2 | 15 | 0 | 0 | 0 | 1 | 7 | 0 | 1 |
Figure
(6 males, 8 females).
Holotype, male, Mount Augustus, WA (
Allotype, female, Yampire Gorge, WA (
Male S3 and S4 broadly emarginated each with round patch of black bristles. Female paraclypeal and supraclypeal area with pale yellow marks, tibial scopa orange, T2 with some black hairs in anterolateral corners, process on S6 irregularly striate, not well defined posteriorly (Fig.
Male holotype (WAM5476): Body length 13 mm, forewing length 9 mm, head width 4.4 mm.
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No. of records: | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 11 | 3 | 0 | 0 | 0 |
Figure
The specific epithet refers to the orange habitus of the species.
3 males, 2 females.
Holotype, male, 7 km NW Barkly Roadhouse, NT (
Allotype, female, 29 km South of Tennant Creek, NT (
Paratype, male WAM 5473, female WAM 5472, same locality data as allotype.
Male with rectangular patch of pale bristles on S4 and shallow emarginated apicomedial area of S5. Female small paraclypeal mark in lower corners, supraclypeal mark present, tibial scopa grey-white, T2 without black hairs anteriorly, process on S6 openly punctate, little defined posteriorly (Fig.
Male holotype (AM K361599): Body length 14 mm, forewing length 9 mm, head width 4.9 mm.
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No. of records: | 0 | 0 | 0 | 0 | 4 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Figure
The specific epithet refers to Michael Batley, the collector of the type specimen, in honour of his contribution to Australian bee taxonomy and identification services.
149 males, 340 females
Lectotype of
Holotype of
Holotype of
Holotype of
Holotype of
Holotype of
Holotype of
Both sexes with orange iridescent metasomal pubescence. Male with small triangular patch of black bristles on S4 and wide triangular emarginated apicomedial area of S5. Female paraclypeal and supraclypeal marks absent, tibial scopa orange, T2 with black hairs anteriorly, process on S6 reticular striate, broadly defined posteriorly (Fig.
Male (AM 361432): Body length 12.5 mm, forewing length 9.7 mm, head width 4.7 mm.
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No. of records: | 53 | 66 | 168 | 112 | 6 | 2 | 2 | 2 | 3 | 5 | 12 | 21 |
Predominantly East coast, Figure
55 males, 49 females
Holotype of
Holotype of
Male with small round patch of black bristles on S4 and triangular emarginated apicomedial area of S5. Female paraclypeal mark absent, supraclypeal mark present, tibial scopa white, T2 without black hairs anteriorly, process on S6 irregular striate and punctate, not defined posteriorly (Fig.
Male holotype
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No. of records: | 2 | 0 | 0 | 0 | 4 | 1 | 0 | 24 | 27 | 26 | 8 | 4 |
When
(18 males, 15 females).
Holotype, male, Wongalara Station, NT (
Allotype, female, same locality data as holotype, SAMA RL2122.
Both sexes body pubescence mostly orange. Male with small round patch of orange bristles on S4, a much smaller patch on S3 and a deep parallel sided emargination on apicomedial area of S5. Female paraclypeal mark absent, supraclypeal mark present, tibial scopa orange, T2 without black hairs anteriorly, process on S6 broadly parabolic and well defined shiny transverse lineo-reticulate and openly punctate (Fig.
Male holotype (SAMA 32-002684, RLb2121): Body length 13 mm, forewing length 10.1 mm, head width 4.7 mm.
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No. of records: | 1 | 1 | 0 | 3 | 9 | 6 | 5 | 1 | 3 | 0 | 3 | 1 |
Figure
The specific epithet refers to the deep parallel medial incision in male S5.
(180 males, 164 females).
Holotype of
The largest species. Sexes are colour-dimorphic: females head and mesosomal pubescence white, males light brown. Body sizes of males have a bimodal distribution, the largest males approximately as large as the females, small males circa ¾ the size. Male with round patch of dark bristles on S4, S5 with apicomedial emargination wide, parabolic. Female paraclypeal and supraclypeal mark present, tibial scopa black, process on S6 small irregular roughened, not well defined (Fig.
Male (VM Holotype T-11869): Body length 16 mm, forewing length 12.4 mm, head width 5.3 mm.
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No. of records: | 0 | 0 | 0 | 1 | 1 | 0 | 42 | 253 | 40 | 1 | 0 | 0 |
The ecology of the mating behaviour of this species has been studied in detail and is summarised in the introduction.
Figure
(2 males, 5 females).
Holotype of
The description of the female holotype of this species can be found in
Pubescence black on T1-T3 (females) and T1-T4 (males), remaining terga white.
Male allotype ANIC Split Rock Qld. Note: Probably mislabelled, likely to be Ubir Rock NT: Body length 17 mm, forewing length 11.1 mm, head width 5.1 mm.
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No. of records: | 2 | 3 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
The collection site Split Rock, Qld of the two males may be a labelling error, as it is outside the distribution of the species as we currently understand it.
no data.
Figure
23 males, 32 females
Holotype of
Holotype of
Holotype of
Examination of the holotype of
Male with rectangular patch of light brown bristles on S4 and emarginated apicomedial area of S5 circa as wide as deep. Female paraclypeal and supraclypeal marks absent, tibial scopa pale yellow or ochre, T2 without black hairs anteriorly, process on S6 large roughly striate and punctate, not well defined posteriorly (Fig.
Male (WAM 5391): Body length 11.8 mm, forewing length 8.8 mm, head width 4.7 mm.
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No. of records: | 16 | 7 | 5 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 4 | 4 |
The type of
Southern temperate Australia, Figure
(4 males, 3 females).
Holotype of
Male with round patch of black bristles on S4 and shallow emarginated apicomedial area of S5. Female paraclypeal and supraclypeal marks present, tibial scopa grey, T2 with band of black hairs anteriorly, process on S6 small narrow transverse lineo-reticulate, shiny posteriorly, not well defined (Fig.
The description of
Male (QM T238472): Body length 14 mm, forewing length 9.3 mm, head width 4.5 mm.
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No. of records: | 1 | 0 | 3 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 |
Figure
(5 males, 7 females).
Holotype, male, Henbury Station, Gloaming Dam, NT (
Allotype, female, Henbury Station, NT (
Male with large patch of dark brown stiff bristles on S4 and emarginated apicomedial area of S5 surrounded by branched hairs. Female paraclypeal and supraclypeal marks present, tibial scopa white, T2-T4 with black hairs anteriorly, process on S6 lineo-reticulate and well defined posteriorly (Fig.
Male holotype (SAMA 32-033609, RL 2222: Body length 16 mm, forewing length 10.3 mm, head width 4.9 mm.
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No. of records: | 0 | 0 | 1 | 8 | 3 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Figure
The specific epithet refers to the greyish hair bands on the metasomal terga.
(1 female).
Holotype of
The description of the female holotype of this species can be found in
Metasomal terga with black pubescence and white posterior hair bands, narrow on T1 (Fig.
A-E.
Month: | Jan | Feb | Mar | Apr | May | Jun | Jul | Aug | Sep | Oct | Nov | Dec |
No. of records: | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Figure
(1 male).
Holotype, male, 55 mls South of Onslow, WA (
Male with large round patch of dark brown stiff bristles on S4 and emarginated apicomedial area of S5. T2-T4 anteriorly with black hairs. Female unknown.
Male (WAM 26816): Body length 14 mm, forewing length 9.2 mm, head width 4.4 mm.
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No. of records: | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 |
This single male specimen from Onslow, WA in the WA Museum collection has an identification label by Brooks with identification ‘
Figure
The specific epithet refers to:
(9 males, 6 females):
Holotype male, Laura Roadhouse, Qld (
Allotype, female, same locality data as holotype, QM T238478;
Paratypes, 1 male, same locality data as holotype, SAMA KR06079; 1 female, Laura Roadhouse, Qld (
Both sexes metasomal terga with black pubescence and light posterior hair bands, sterna black and shiny. Male with two small adjacent patches of black bristles on S4 and two large comma-shaped patches of bristles on S5. Female paraclypeal marks absent and supraclypeal marks present, tibial scopa black with white dorsal streak, process on S6 well defined with smooth posteriorly protruding lobe (Fig.
Male holotype (QM T238477, RLa2427): Body length 12 mm, forewing length 9.2 mm, head width 4.3 mm.
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No. of records: | 0 | 0 | 15 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Figure
The specific epithet refers to shiny black metasomal sterna in both sexes.
45 males, 57 females.
Holotype of
Holotype of
Holotype of
Holotype of
Examination and comparison of the female types of
Male with rectangular patch of dark and light brown bristles on S4 and angular emarginated apicomedial area of S5 surrounded by branched hairs. Female paraclypeal and supraclypeal marks absent, tibial scopa grey-white, T2 with narrow band of black hairs anteriorly, process on S6 smooth and well defined posteriorly (Fig.
Male (VM HYM-35946): Body length 13.5 mm, forewing length 9.7 mm, head width 5.1 mm.
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No. of records: | 3 | 54 | 27 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
When
Figure
(32 males, 22 females).
Syntypes of
Holotype of
Examination of the male holotype of
Both sexes metasomal integument reddish. Male with small round patch of orange bristles on S4, a much smaller patch on S3 and a deep (circa half the width of the segment) emargination on apicomedial area of S5. Female paraclypeal marks absent and supraclypeal marks small, tibial scopa orange, terga lacking black hairs anteriorly, process on S6 well defined broad parabolic and lineo-reticulate (Fig.
Male (WAM 5469): Body length 13 mm, forewing length 10.4 mm, head width 5 mm.
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No. of records: | 7 | 7 | 5 | 2 | 1 | 3 | 1 | 2 | 0 | 2 | 4 | 10 |
Figure
(45 males, 66 females).
Holotype, male, Beltana Station, SA (30.6179S: 139.3214E), 15 Jun. 2009, R & P Leijs, on
Allotype, female, Andamooka Homestead, SA (
Paratypes, 3 females, Port Augusta Arid Lands Botanic Garden, SA (
Male with wide rectangular patch of brown bristles on S4 and widely emarginated apicomedial area of S5, apex of S6 with conspicuous two-coloured y-shaped hair brush. Female paraclypeal marks absent, supraclypeal marks present, tibial scopa pale orange, T2 with band of black hairs anteriorly, process on S6 large transverse lineo-reticulate, not well defined posteriorly (Fig.
Male (SAMA 32-033613, RL1452): Body length 15 mm, forewing length 11 mm, head width 4.9 mm.
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No. of records: | 0 | 0 | 0 | 0 | 3 | 8 | 17 | 48 | 24 | 2 | 1 | 0 |
Some field observations on mating behaviour and nests can be found on the following link:
Central Australia, the northern limit of the distributions is just north of the tropic of Capricorn, Figure
The specific epithet refers to the wide brush of stiff setae apicomedially on S4 of the male.
Holotype of
Female paraclypeal and supraclypeal marks absent, tibial scopa pale orange, T2 without band of black hairs anteriorly. Scutum and terga pubescence uniformly ochre brown.
This specimen could not be matched with any other species, or any of the specimens examined. Based on the diagnostic characters mentioned above it would key out to
(11 males, 1 female).
Holotype, male, 19 km SE of Laura, Qld (
Allotype, female, Iron Range, Cape York Qld (
Paratypes, 3 males, same locality data as holotype, QM T238475, SAMA KR06115, SAMA KR06118, 1 male 6.7 km NE of Laura, Qld (
Both sexes metasomal terga entirely with black pubescence.
Male holotype (QM T238476, RL2433b): Body length 16 mm, forewing length 12.7 mm, head width 5.6 mm.
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No. of records: | 0 | 0 | 7 | 0 | 0 | 4 | 0 | 0 | 0 | 0 | 0 | 0 |
At three different locations in 2017, male territorial behaviour was observed above water with native Water Hyacinth,
Figure
The specific epithet refers to Australian female carpenter bees
(1 female):
Holotype, female, Litchfield, NT (
Male unknown. Female T2-T4 with black pubescence and pale posterior hairbands, T5 entirely with white hairs, clypeus with two black sub-parallel marks, process on S6 smooth with well-developed broad ridge (Fig.
Female holotype (SAMA 32-033601, RL707): Body length 13 mm, forewing length 9.1 mm, head width 4.5 mm.
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No. of records: | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
None.
Figure
The specific epithet refers to Andy Young, the collector of the specimen, in honour of his contribution to
This paper would not have been completed without the support of the Bush Blitz program provided through ABRS. We thank Jo Harding, Kate Gillespie, Mim Jambrecina, Beth Tully, Brian Hawkins, and the other members of the Bush Blitz team for their fabulous support in the field. We thank Susan Wright (QM), Nicole Fisher (ANIC), Peter Hudson (SAMA), Brian Hanich (WAM), and Terry Houston (WAM) for providing access to their collections. We thank Wiebe Hogendoorn for advice and checking the new scientific names and Michael Orr for valuable comments on the manuscript. This work was supported by a Bush Blitz Tactical Taxonomy Grant from the Australian Government through ABRS.
Supplementary table - examined specimen data
Microsoft Excell Spreadsheet (.xlsx).
Examined specimen data.