First records of the genera Sivaloka Distant, 1906, with two new species from China, and description of a new species of genus Kodaianella Fennah, 1956 (Hemiptera, Fulgoromorpha, Issidae)

Abstract The genus Sivaloka Distant, 1906 (Hemisphaeriinae, Kodaianellini) is recorded from China for the first time, with two new species Sivaloka arcuata Chang & Chen, sp. nov. (China: Guizhou) and Sivaloka trigona Chang & Chen, sp. nov. (China: Guangxi). One new species of Kodaianella Fennah, 1956, Kodaianella furcata Chang & Chen, sp. nov. (China: Guangxi) is also described and illustrated; female genitalia of two known species in Kodaianella are described. A checklist of species of the tribe Kodaianellini with their distribution and a key to genera are provided.

Female genitalia . Anal tube (Figs 17, 23) oblong, obviously longer in middle than its width; apical margin arched, convex; lateral margin parallel. Anal style (Figs 17, 23) long or short, located near base of anal tube. Hind margin of gonocoxa VIII with endogonocoxal lobe not obvious (Figs 18,24); endogonocoxal process membranous, gradually narrowing. Anterior connective lamina of gonapophyses VIII (Figs 18,24) irregularly rectangular, with two lateral teeth bearing two keels in lateral group and three teeth in apical group. Posterior connective lamina of gonapophyses IX (Figs 19,20,25,26) triangular, with lateral field membranous; sublateral field with two sclerous triangular processes in lateral margin (Figs 19,25); median field with unobvious prominence (median dorsal process) (Figs 19,25); ventroposterior lobes acutely bent at an angle (posterior ventral lobes) (Figs 20,26 Anal tube with apical margin nearly truncated near middle part (Fig. 9); aedeagus with pair of biforked long hooks in ventral view (Fig. 13) Zhang & Chen, 2010 in appearance, but it differs from the latter in having the phallobase with dorsal lobe bearing one rod-like process near its apical part in lateral view (Fig. 12a); the phallobase with ventral lobe distinct short, apical part finger-liked in ventral view (Fig. 13c); and the aedeagus in lateral view, with a forked and hooked process near the apical 1/3 (Fig. 12d).
Etymology. The new species is derived from the Latin word "furcata", in reference to the aedeagus, which bears a forked, hooked process.

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Phallobase with dorsal lobe with small spinous process near apical part (Fig. 54a); aedeagus with one hooked process near basal 1/3 in lateral view, directed to cephalad (Fig. 54e)  Diagnosis. This species is identified by the dark-brown or yellow-brown frons, without any bands (Fig. 33); the clypeus without median carina, with its basal part swollen and its apical part sunken (Fig. 33); the forewings longer 2.0 times than their width (Figs 30, 35); the dorsal margin of the genital styles bearing one arched prominence near the middle (Fig. 40); the phallobase with the dorsal lobe bearing a relatively long, hooked process near its apical part (Fig. 42a) and with a triangular process on the ventral margin (Fig. 42b); and the aedeagus near its middle in lateral view with one hooked process, which is directed to caudad (Fig. 42e).
Male genitalia. Anal tube in dorsal view (Fig. 39) longer in middle than the widest breadth (2.50: 1.00), the maximum width in middle of anal tube; apical margin distinctly, angularly concave; lateral margins almost parallel, slightly concave near apical 2/3 of anal tube. Anal style (Fig. 39) relatively long and stout, located in basal 2/5 of anal tube, not surpassing the opening of anal pore. Pygofer (Fig. 38) irregularly rectangular; anterior and posterior margins nearly parallel in lateral view; dorsal and ventral margins parallel. Genital styles (Fig. 40) irregularly triangular in lateral view; dorsal and ventral margins not parallel; dorsal margin with one arched prominence near middle at base of capitulum; ventral margin slightly arched. Capitulum of genital styles irregularly triangular, with small lobe; with stout and not obvious neck (Fig. 41). Phallobase (Figs 42,43) with dorsal lobe slightly expanded into membranous, cystiform process; and dorso-lateral lobe splitting into relatively long, hooked process near apical part (Figs 42a, 43a), ventral margin of dorso-lateral lobe with angular process in basal 1/3 in lateral view (Fig. 42b); lateral lobe distinctly shorter than dorsal lobe in lateral view (Fig. 42c), splitting into two stout branches (Fig. 43c); ventral lobe relatively longer than lateral lobe in lateral view (Fig. 42d) in lateral view; in ventral view, apical part obviously archedly convex (Fig. 43d); lateral margins parallel in ventral view. Aedeagus (Figs 42,43) with one long, hooked process near middle (Fig. 42e) in lateral view, directed to caudad.
Etymology. The specific name is derived from the Latin words "arcuata" in reference to the genital styles which bear an arched prominence on the base before the capitulum.
Description. Body length (from apex of vertex to tip of forewings): male 5.00 mm; forewings: male 4.10 mm.
Etymology. The specific name is derived from the Latin words "trigona" in reference to the triangular prominence near the middle of the dorsal margin of the genital styles.

Discussion
A comparison of Kodaianella Fennah, 1956, Sivaloka Distant, 1906, and Dentatissus Chen, Zhang & Chang, 2014 shows that species in these genera look rather similar. In these genera the width of the vertex at the base is longer than its length at its middle, the frons lacks transverse carina, the hingwings have A 11 unbranched, and the male genitalia are in general similar.
Sivaloka is, however, clearly different from the other two genera in having the Pcu and A 1 veins on the forewings keel-shaped ( Fig. 2; Chen et al. 2014: fig. 2-79B). There are also significant generic differences in the structure of the male genitalia among the three genera. In Sivaloka, the anal tube is irregularly rectangular with its lateral margins nearly parallel (Figs 39, 51), while in Kodaianella the anal tube is irregularly triangular with the lateral margin apically diverging (Fig. 9); in Dentatissus the anal tube is oval and wider near its middle (Chen et al. 2014: fig. 2-79H). The capitulum of the genital styles are with a stout irregular triangular at their base in Dentatissus (Chen et al. 2014: fig. 2-79J); the capitulum are absent Kodaianella and Sivaloka (Figs 10, 40). The dorsal lobe of the phallobase bears various processes near its apical part in Kodaianella and Sivaloka (Figs 12, 42), but these processes are absent in Dentatissus. The aedeagus bears two pairs of hook-like processes in Dentatissus (Chen et al. 2014: fig. 2-79J) and one pair of hook-like processes in Kodaianella and Sivaloka.