The ML tree (ln L = −23350.60) for 5,124 bp of nuclear 18S rDNA and histone H3, and mitochondrial COI, tRNACys, tRNAMet, 12S rDNA, tRNAVal, 16S rDNA, and ND1 markers. A species name of Orobdella in red indicates a quadrannulate species; in green, sexannulate; and in blue, octannulate. The numbers associated with the nodes represent the bootstrap values for ML (BS)/and Bayesian posterior probabilities (BPPs).

 
 
  Part of: Nakano T (2014) A new quadrannulate species of Orobdella (Hirudinida, Arhynchobdellida, Orobdellidae) from central Honshu, Japan. ZooKeys 445: 57-76. https://doi.org/10.3897/zookeys.445.7999