Pheidole makaensis sp. nov., head and profile of major worker (A), hypostomal teeth (J), profile of minor worker (F). Pheidole fitarata sp. nov., head and profile of major worker (B), hypostomal teeth (K), profile of minor worker (G). Pheidole rugofitarata sp. nov., head and profile of major worker (C), hypostomal teeth (L). Pheidole avaratra sp. nov., head and profile of major worker (D), hypostomal teeth (M), profile of minor worker (H). Pheidole ehazoara sp. nov., head and profile of major worker (E), hypostomal teeth (N), profile of minor worker (I).