Porongurup, a new genus of pselaphine staphylinid beetles from Western Australia (Coleoptera, Staphylinidae, Pselaphinae, Faronitae)

Abstract A new genus and three new species of pselaphine staphylinid beetles, supertribe Faronitae, from Western Australia are described as follows: Porongurupgen. nov. is based on Porongurup angulatussp. nov., with the two additional new species, Porongurup clarkeisp. nov. and Porongurup tenuissp. nov. Illustrations of their habitus, and major diagnostic characters as well as a distribution map are included. A key to the species is provided.


Introduction
The supertribe Faronitae consists of 29 genera worldwide, with 13 genera being found in Australia and New Zealand (Park and Chandler 2017). At this time only three faronite genera [Sagola Sharp, 1874 (9 spp.), Logasa Chandler, 2001 (3 spp.), and Nornalup Park & Chandler, 2017 (3 spp.)] are known from Australia, although eastern Australia does have a complex and largely unknown fauna of Faronitae that is currently under study by J.-S. Park and colleagues. This is the second paper treating the separate and unique fauna of Faronitae from the southwestern corner of Australia, following the treatment of Nornalup by Park and Chandler (2017).
In the collections of Faronitae being examined as part of a treatment of the fauna of Australia, 31 specimens were segregated into three species that shared a combination of discrete diagnostic characters, with that combination separating them from all other known faronite genera and supporting the creation of a new genus. This new genus can be recognized by the following combination of characters: rostrum with linear frontal sulcus, abdominal segment VI approximately twice as long as V, and the male genitalia have the median lobe of the male genitalia elongate and narrow, culminating in an acute apex. The included species of this genus are known only from the southwestern corner of Western Australia.
The two faronite genera known from extreme southwestern Australia, Porongurup gen. nov. and Nornalup, are apparently restricted to this area, which is known as a global biodiversity hotspot that has a climatically isolated flora and fauna (Slater 1975, Coates and Atkins 2001, Hopper and Gioia 2004, Slatyer et al. 2007, Park and Chandler 2017. Members of Porongurup gen. nov. and Nornalup have been collected together in samples from this area, and apparently share the same microhabitats. However, the species of Porongurup have never been recorded together in samples, indicating differences in habitat preferences for these species.

Materials and methods
Thirty-one specimens were examined from the following collections: Field Museum of Natural History (FMNH), Chicago, Illinois, USA, and the University of New Hampshire Insect Collection (UNHC), Durham, New Hampshire, USA. Six specimens were mounted on permanent slides to aide in observation of the internal characters and the fine external characters that are not apparent when using a dissecting microscope. Permanent microscopic slides were prepared using the techniques described by Hanley and Ashe (2003). Terminology for the foveal system and nomenclature follows Chandler (2001). Decimal Degrees were used for the format of geographical coordinates. Holotypes are deposited in the Western Australian Museum (WAM), Perth, Western Australia, Australia, and paratypes are deposited in the Field Museum of Natural History, the Western Australian Museum, the Australian National Insect Collection (ANIC), Canberra, ACT, Australia, the University of New Hampshire Insect Collection, Durham, New Hampshire, USA, and the Chungbuk National University Insect Collection (CBNUIC), Cheongju, Chungbuk-do, South Korea (indicated parenthetically). Specimen label data for the holotypes is transcribed verbatim. Data from the paratypes are standardized for consistency. The map of Australia is based on an image from SimpleMappr (Shorthouse 2010), that was subsequently modified to indicate localities of the specimens.

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Antennomere III subquadrate, as long as wide (Fig. 4A, B); female elytra longer than wide with fully developed hind-wings (Fig. 1A, D); length of abdominal segment VI 2.0-2.5 times as long as V (Fig. 1A, D Diagnosis. This species can be distinguished from Porongurup clarkei sp. nov. by the smaller size of the male genitalia (Fig. 2), the subquadrate antennomere III (Fig. 4A, B), and the length of abdominal segment VI is approximately twice as long as V. This species is also separated from Porongurup tenuis sp. nov. by the length of abdominal segment VI being twice as long as V (Fig. 1), and the parameres of the male genitalia approximately are twice as wide as those of the other species (Fig. 2).  Description. Length 1.7-1.9 mm (Fig. 1A, D). Head. Head in dorsal view with deep frontal sulcus and vertexal foveae (Fig. 4G). Antennomeres II longer than wide, III subquadrate, smallest of the antennomeres, IV and V longer than wide, VI and VIII as long as wide, IX and X transverse (Fig. 4A, B). Abdomen. Length of abdominal segment VI twice as long as V in both sexes (Fig. 1A, D). Aedeagus. Median lobe of male genitalia elongated triangular, lacking articulated extension at tip of apex. Phallobase rounded in dorsal view. Parameres as wide as at middle of median lobe ( Fig. 2A).
Etymology. This species name refers to the sub-apically angulate parameres of the male genitalia.
Habitat. Specimens of this species were collected using flight intercept traps, or were taken by sifting leaf, bark, log, or fungus in Eucalyptus forests. Diagnosis. This species can be distinguished from Porongurup angulatus sp. nov. by antennomere 3 being longer than wide (Fig. 4C), and abdominal segment VI being approximately three times longer than V (Fig. 1B, E). This species is also separated from Porongurup tenuis sp. nov. by the median lobe of the male genitalia having a small digit at the apex ( Fig. 2B: arrowed).
Description. Length 1.6-1.9 mm (Fig. 1B, E). Head. Head in dorsal view with both shallow frontal sulcus and vertexal foveae. Male and female antennae are almost identical in length (Fig. 4C, D). Antennomeres II longer than wide, III subquadrate and smallest, IV-VIII longer than wide, IX and X transverse (Fig. 4C, D). Elytra. Male elytra longer than wide, female elytra shorter, as long as wide (Fig. 1B, E). Abdomen. Abdominal segment VI approximately three times longer than V (Fig. 1B, E). Aedeagus. Median lobe of male genitalia as long as parameres with apical articulated extension. Phallobase rounded in dorsal view. Parameres symmetrical (Fig. 2B, C).
Etymology. This species is named for one of the collectors of the holotype, the staphylinid specialist Dave J. Clarke.
Habitat. Specimens of this species were collected by sifting leaf, log, bark, moss or fungus litter in Eucalyptus forests, or were taken by Berlese funnel water-washed soil. Diagnosis. This species can be distinguished from Porongurup angulatus sp. nov. by the elytra being as long as wide (Fig. 1C, F). It also differs from P. clarkei sp. nov. by lacking the articulated extension at the apex of the median lobe of the male genitalia (Fig. 2D).
Description. Length 1.8-2.0 mm (Fig. 1C, F). Head. Head in dorsal view with both deep and narrow frontal sulcus and vertexal foveae. Male antennae longer than those of female (Fig. 4E, F). Antennomeres II longer than wide, III subquadrate and smallest, IV and V longer than wide, VI-VIII as long as wide, IX and X transverse (Fig. 4E, F). Elytra. Male elytra longer than wide, female elytra shorter than those of male (Fig. 1C, F). Abdomen. Male abdominal segment VI approximately three times as long as V, female with segment VI 1.5 times longer than V (Fig. 1C, F). Aedeagus. Median lobe of male genitalia as long as parameres, apex sharp and narrow. Phallobase oval in dorsal view. Parameres symmetrical (Fig. 2D).
Etymology. This species name refers to the elongate slender parameres of the male genitalia.
Habitat. Specimens of this species were collected using flight intercept traps, or by sifting leaf or log litter.