Mario Cupello, Cibele S Ribeiro-Costa, Fernando Z Vaz-De-Mello (2022)
The evolution of Bolbites onitoides (Coleoptera: Scarabaeidae: Phanaeini): its phylogenetic significance, geographical polychromatism and the subspecies problem.
Zoological Journal of the Linnean Society194: 973.
DOI: 10.1093/zoolinnean/zlab015