Map showing Costa Rican scarabaeine endemic numbers as distributed by life-zones (taken from Kohlmann et al. 2007).

 
 
  Part of: Kohlmann B, Solís Á, Alvarado G (2019) Description of Onthophagus humboldti and Uroxys bonplandi, two new scarab beetles (Coleoptera, Scarabaeidae, Scarabaeinae) from Costa Rica, with notes on tropical mountain brachyptery and endemicity. ZooKeys 881: 23-51. https://doi.org/10.3897/zookeys.881.38026