Maps of Madagascar with species distributions and administrative divisions. A Copelatus amphibius sp. nov. (circle), Copelatus betampona sp. nov. (square) B Copelatus zanatanensis sp. nov. (circle), Madaglymbus semifactus sp. nov. (square) C Madaglymbus kelimaso sp. nov. (circle), Madaglymbus menalamba sp. nov. (square) D current 22 regions and six former provinces of Madagascar.

 
 
  Part of: Ranarilalatiana T, Bergsten J (2019) Discovery of a specialist Copelatinae fauna on Madagascar: highly ephemeral tropical forest floor depressions as an overlooked habitat for diving beetles (Coleoptera, Dytiscidae). ZooKeys 871: 89-118. https://doi.org/10.3897/zookeys.871.36337