Male genitalia. From left to right aedeagus in right lateral, ventral, left lateral views, and left paramere. A Copelatus amphibius sp. nov. B Copelatus zanatanensis sp. nov. C Copelatus betampona sp. nov.

 
 
  Part of: Ranarilalatiana T, Bergsten J (2019) Discovery of a specialist Copelatinae fauna on Madagascar: highly ephemeral tropical forest floor depressions as an overlooked habitat for diving beetles (Coleoptera, Dytiscidae). ZooKeys 871: 89-118. https://doi.org/10.3897/zookeys.871.36337