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  • ContentsContents
  • Article InfoArticle Info
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  • Article title
  • Abstract
  • Keywords
  • Introduction
  • Materials and methods
    • Fieldwork
    • Preparations and illustrations
    • Material and depositories
  • Taxonomy
    • Copelatus
    • Copelatus amphibius
    • Copelatus zanatanensis
    • Copelatus betampona
    • Madaglymbus
    • Madaglymbus semifactus
    • Madaglymbus kelimaso
    • Madaglymbus menalamba
  • Discussion
  • Acknowledgements
  • References
  • Supplementary material
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