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Article title
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Abstract
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Keywords
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Introduction
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Materials and methods
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Fieldwork
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Preparations and illustrations
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Material and depositories
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Taxonomy
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Copelatus
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Copelatus amphibius
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Copelatus zanatanensis
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Copelatus betampona
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Madaglymbus
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Madaglymbus semifactus
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Madaglymbus kelimaso
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Madaglymbus menalamba
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Discussion
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Acknowledgements
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References
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Supplementary material