Corresponding author: Qiang Yang (
Academic editor: Shaun Winterton
A new genus and species of
Yang Q, Shi C, Ren D (2019) A new genus and species of berothids (Insecta, Neuroptera) from the Late Cretaceous Myanmar amber. ZooKeys 864: 99–109.
List of named fossil
Species | Age | Locality | Reference |
---|---|---|---|
Middle Jurassic Bathonian to Callovian | Inner Mongolia, China (Jiulongshan Formation) |
|
|
Middle Jurassic Bathonian to Callovian | Inner Mongolia, China (Jiulongshan Formation) |
|
|
Middle Jurassic Bathonian to Callovian | Inner Mongolia, China (Jiulongshan Formation) |
|
|
Upper Jurassic Upper Callovian–Kimmeridgian | Karatau, Kazakhstan (Karabastau Formation) |
|
|
Upper Jurassic Upper Callovian–Kimmeridgian | Karatau, Kazakhstan (Karabastau Formation) |
|
|
Upper Jurassic Upper Callovian–Kimmeridgian | Karatau, Kazakhstan (Karabastau Formation) |
|
|
Upper Jurassic Upper Callovian–Kimmeridgian | Karatau, Kazakhstan (Karabastau Formation) |
|
|
Upper Jurassic Upper Callovian–Kimmeridgian | Karatau, Kazakhstan (Karabastau Formation) |
|
|
Upper Jurassic Upper Callovian–Kimmeridgian | Karatau, Kazakhstan (Karabastau Formation) |
|
|
Upper Jurassic | Khoutiyn-Khotgor, Mongolia (Ulan-Ereg Formation) |
|
|
Early Cretaceous Early Berriasian | Durlston Bay, England (Lulworth Formation) |
|
|
Early Cretaceous Valanginian/Hauterivian | Lebanese amber (Jezzine) |
|
|
Early Cretaceous Valanginian/Hauterivian | Lebanese amber (Jezzine) |
|
|
Early Cretaceous Barremian | Liaoning, China (Yixian Formation) |
|
|
Late Cretaceous lowermost Cenomanian | Myanmar amber | This paper | |
Late Cretaceous lowermost Cenomanian | Myanmar amber |
|
|
Late Cretaceous lowermost Cenomanian | Myanmar amber |
|
|
Late Cretaceous lowermost Cenomanian | Myanmar amber |
|
|
Late Cretaceous lowermost Cenomanian | Myanmar amber |
|
|
Late Cretaceous lowermost Cenomanian | Myanmar amber |
|
|
Late Cretaceous lowermost Cenomanian | Myanmar amber |
|
|
Late Cretaceous lowermost Cenomanian | Myanmar amber |
|
|
Late Cretaceous lowermost Cenomanian | Myanmar amber |
|
|
Late Cretaceous lowermost Cenomanian | Myanmar amber |
|
|
Late Cretaceous lowermost Cenomanian | Myanmar amber |
|
|
Late Cretaceous lowermost Cenomanian | Myanmar amber |
|
|
Late Cretaceous lowermost Cenomanian | Myanmar amber |
|
|
Late Cretaceous lowermost Cenomanian | Myanmar amber |
|
|
Late Cretaceous Turonian | Raritan (New Jersey) amber |
|
|
Late Cretaceous Turonian | Raritan (New Jersey) amber |
|
|
Late Cretaceous Turonian | Raritan (New Jersey) amber |
|
|
Early Eocene | Hat Creek amber, British Columbia |
|
|
Late Eocene | Baltic amber |
|
|
Early Eocene late Ypresian | Colorado, USA (Green River Formation) |
|
This study is based on one female specimen from Myanmar amber. The amber pieces were collected in the Hukawng Valley (the state of Kachin in northern Myanmar). A detailed map of the Hukawng Valley is given by
Venational terminology generally follows
Abbreviations:
The generic name is a combination of the Latin
Antenna long, more than 6.6 mm, longer than body or forewings; scape elongate, ca. 0.64 mm, almost 12 times as long as wide; flagellum with about 100 flagellomeres. Pronotum elongate, about three times as long as wide. Forewing with one basal sc-r and four ra-rp, M forked distal to the separation of
The specific epithet is named after Ms Jiewen Zhao (Hunan, China), the daughter of this amber’s owner (Ms Dan Zuo). Her mother hopes that this honour will promote Jiewen’s interests in natural history.
As for the genus.
CNU-NEU-MA2018072, female, a nearly complete and well-preserved specimen.
Hukawng Valley, Kachin State, northern Myanmar; lowermost Cenomanian, Upper Cretaceous.
Holotype CNU-NEU-MA2018072. Total body length 4.0 mm. Head and body with numerous scattered, fine setae; head about as wide as long. Compound eyes large. Antenna filiform, over 6.6 mm, with scattered setae all over; scape elongate, ca. 0.64 mm, almost 12 times as long as wide; pedicel as long as wide, slightly thicker than flagellum; flagellum with approximately 100 flagellomeres, the last few flagellomeres tapering. Pronotum elongate, narrower than head, about three times as long as wide; pro-, meso-, and metanotum with scattered, long, fine setae. Legs relatively long and slender, with numerous short setae intermixed with long setae. Forelegs: coxa elongated; femur long and slender; tibia slightly inflated nearly as long as femur; basitarsus nearly three times as long as the second tarsomere, the last four tarsomeres of the same length, each tarsomeres with two ended spur. Mid- and hind legs coxa coniform, thicker than forelegs. Each leg with two pretarsal claws, one big arolium. Abdomen nine segments, with scattered short setae; gonapophysis lateralis elongate.
Forewing length 5.5 mm, width 1.5 mm (left forewing/LFW); length 4.9 mm, width 1.8 mm (right forewing/RFW); elongated ovoid, apex rounded, with dense relatively short setae on veins and longer setae on margins; trichosors prominent along entire wing margin. Humeral vein crossvein-like; presumable
Hind wing elongate, length 5.1 mm, width 1.5 mm (left hind wing/LHW); length 5.2 mm, width 1.5 mm (right hind wing/RHW). Trichosors prominent along entire wing margin. Costal space narrow, dilated distal to the fusion of
We appreciate the valuable comments and useful suggestions on our manuscript from the editor (Dr. Shaun L. Winterton), the reviewer (Dr. Vladimir N. Makarkin), and another anonymous reviewer. We thank Ms Dan Zuo (Changsha, Hunan, China) for donated of the type specimen to us for study. This study was supported by National Natural Science Foundation of China (grant nos. 41602014, 31501881, 31730087), Program for Changjiang Scholars and Innovative Research Team in University (grant no. IRT-17R75), Project of High-level Teachers in Beijing Municipal Universities (IDHT20180518), Scientific Research Foundation of Guangzhou University (69-18ZX10150).