A simplified species-level phylogeny derived from the BI tree in the parallel paper (Xu et al. 2019) showing three species groups.

 
 
  Part of: Xu X, Ono H, Kuntner M, Liu F, Li D (2019) A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50. https://doi.org/10.3897/zookeys.888.34494