Male genitalia. From left to right, aedeagus in right lateral, ventral, left lateral views and left paramere. A Copelatus insuetus B Copelatus kely sp. nov. C Copelatus vokoka sp. nov. D Copelatus ankaratra sp. nov. E Copelatus safiotra sp. nov.

 
 
  Part of: Ranarilalatiana T, Raveloson Ravaomanarivo LH, Bergsten J (2019) Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species. ZooKeys 869: 19-90. https://doi.org/10.3897/zookeys.869.33997