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  • Article title
  • Abstract
  • Keywords
  • Introduction
  • Materials and methods
    • Type revision, morphological terminology, and classification
    • Geological settings
    • Geometric morphometric analyses of forewing shape
    • Shape discrimination at different taxonomic levels
    • Assignment of the bee fossils
  • Results
    • Geometric morphometric analyses
    • Systematics
    • Family Apidae Latreille
      • Late Eocene
      • Eocene-Oligocene boundary
      • Oligocene-Miocene boundary
      • Lower Miocene
      • Middle-Lower Miocene
      • Upper Miocene
  • Discussion
    • Geometric morphometrics of forewing shape to discriminate taxa
    • Taxonomic affinities of Calyptapis florissantensis and Oligobombus cuspidatus
    • Origin and diversification of bumble bees
    • Cenozoic extinctions of Corbiculata
  • Acknowledgements
  • References
  • Supplementary materials
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