Synopsis of Acanthocerini (Hemiptera, Coreidae) from Argentina

Abstract Eight genera and 13 species of the tribe Acanthocerini are recorded in Argentina, i.e., Athaumastus haematicus (Stål), Athaumastus macer Brailovsky, Athaumastus subcarinatus (Stål), Athaumastus subterlineatus Bergroth, Beutelspacoris sanchezi Brailovsky, Beutelspacoris dilatata Casini, Camptischium clavipes (Fabricius), Crinocerus sanctus (Fabricius), Dersagrena flaviventris (Berg), Dersagrena lacerdae (Signoret), Dersagrena subfoveolata (Berg), Thlastocoris hernandezi Brailovsky and Zoreva dentipes Fabricius. Redescriptions are given for Athaumastus haematicus, Athaumastus subcarinatus and Dersagrena flaviventris with photographs of male and female genitalia of Dersagrena subfoveolata. Zoreva recorded from Argentina the first time. New locality records are given for Buenos Aires, Chaco, Formosa, Misiones, and Tucumán.


introduction
The Coreidae, commonly called "leaf-footed, pod-or squash-bugs", are heavy bodied insects usually strongly elongate or broadly elliptical. This family includes some of the largest living heteropterans, as well as species that are delicate or slender (Schuh & Slater 1995). The family contains about 267 genera and more than 1884 species worldwide (Henry 2009). The basic suprageneric classification was established by Stål (1867Stål ( , 1870 and Schaefer (1964Schaefer ( , 1965, who provided the most comprehensive treatment. Packauskas (1994) gave keys to the subfamilies and tribes of the New World Coreidae and a checklist of published keys to genera and species, and Packauskas (2010) cataloged the New World Coreidae and provided a comprehensive introduction to the literature.
The most comprehensive treatment of the Coreidae for Argentina are by Pennington (1920Pennington ( , 1921 and Kormilev (1954), but only Pennington (1921) dealt with the Acanthocerini.
Knowledge of the South American fauna is poor; this is especially true in regard to economically important taxa. Comprehensive keys for identification of the species in the region have not been published.
Most of Argentina lies in the Neotropical faunal region. The country covers an area of 2,791,810 km 2 and is bordered by Uruguay, Brazil, Paraguay, Bolivia, and Chile. Approximately 75% of the country is occupied by arid and semiarid areas, but rainforests are also present in the northeast, i.e., the Yungas and Paranaense regions. Coscarón (submitted) recorded 125 species in 48 genera of Coreidae living from Argentina.
The goal of this paper is to provide an illustrated key to the genera of Acanthocerini (Coreidae) from Argentina (Fig. 38), a diagnosis and redescription of some of its species, geographical distribution, and a list of species for each genus.

Materials and methods
All specimens from this study are deposited in the collections of the Museo de Ciencias Naturales de La Plata (MLP), La Plata, Buenos Aires, Argentina (http://www.fcnym.unlp. edu.ar/abamuse.html). Photographs were compared with material of the Naturhistoriska Riksmuseet in Stockholm, Sweden (http://www.nrm.se/2.1286b10fdbe80efba80001. html) and the American Museum of Natural History in New York (http://www.amnh. org/). For the geographical distribution we used the program DIVA-GIS 7.1.7 (http:// www.diva-gis.org/) and the distribution of those specimens for which global positioning system data were available was used for the construction of maps. Photographs were produced using a Kodak Easy Share (12 megapixels) camera and a magnifying Wild M-Stereomicroscope. The diagnoses of genera are taken from Brailovsky (1987) and O´Shea (1980), except as noted. Coreoidea Species File (Version 1.1/4.1) (http://coreoidea.speciesfile.org/HomePage.aspx) was consulted. All measurement are in millimeters.