Zhezhoulinyphia gen. nov. (Araneae, Linyphiidae) from Yunnan, China

Abstract A new Linyphiinae genus Zhezhoulinyphiagen. nov. from Yunnan Province is described together with its two new species: Z.caperatasp. nov. (♂) and Z.denticulatasp. nov. (♂, ♀). Centromerusyadongensis Hu & Li, 1987 is transferred to Zhezhoulinyphiagen. nov. and a new combination, Z.yadongensis (Hu & Li, 1987), comb. nov. is proposed.


Introduction
Linyphiidae Blackwall, 1859, the second largest spider family, contains 4591 species in 611 genera, which accounts for approximately 10% of the total number of spiders worldwide (WSC 2019). Li and Lin (2016) listed 371 Chinese linyphiid species in 154 genera, including 69 species in 16 genera in the subfamily Linyphiinae. Yunnan is a biodiversity-rich spot in south China and about 9 genera and 39 species of linyphiid spiders have been described from there (Wunderlich and Song 1994;Xia et al. 2001;Liu and Chen 2010;Zhao and Li 2014;Zhao and Li 2017;Irfan and Peng 2018).
While examining the specimens collected from Gaoligong Mountain, Zhezhoulinyphia gen. nov. with two new species were identified and are described here.

Material and methods
Specimens were collected by hand picking and beating shrubs and were kept in 75% ethanol. The type specimens are deposited at the College of Life Sciences, Hunan Normal University, Changsha, China. After dissection, epigyna were cleared in trypsin enzyme solution before examination and photography. The left male palp was used for description and illustration. Specimens were examined, measured with a Leica M205C stereomicroscope. Photos were taken with a digital camera Canon PowerShot G12 mounted on an Olympus BX53 and Leica MC170 HD mounted on a Leica M205C. Stacked focus images were generated using Helicon Focus software (3.10, free). A map was created using ArcMap 10.2, and then modified using Adobe Photoshop CS2 Extended. Leg measurements are given in the following order: total length (femur, patella + tibia, metatarsus, tarsus). All measurements are given in millimeters (mm).
Abbreviations used in the text and figures are as follows: AE anterior wall of epigyne; AER anterior eyes row; ALE anterior lateral eyes; AME anterior median eyes; AME-AME distance between AME; AME-ALE distance between AME and ALE; Type species. Zhezhoulinyphia denticulata sp. nov. Etymology. The species name comes from the Chinese word "褶皱(Zhezhou)" meaning "wrinkle" and referring to parmula with wrinkles in epigyne. Gender feminine.

Discussion
Zhezhoulinyphia gen. nov. can be identified as the member of the subfamily Linyphiinae by the following morphological characters: female palp with claw; male palp without tibial apophysis; maxillae long and parallel; all the tibiae with two dorsal spines (Merrett 1963